Oenothera hartwegii subsp. pubescens
Onagraceae tribe Onagreae
0.6–4 × 0.15–1.2 cm, fascicles of small leaves often absent or much reduced, sometimes to 1.5 cm;
blade very narrowly elliptic or narrowly lanceolate to ovate, base truncate or subcordate and clasping, rarely nearly clasping, margins entire or sparsely serrulate, rarely crinkled-undulate.
alternate or basal;
stipules absent.
buds with free tips 0.5–3 mm;
floral tube 20–50 mm;
sepals 9–26 mm;
petals 12–35 mm;
filaments 5–12 mm, anthers 4–13 mm;
style 25–70 mm.
usually actinomorphic, rarely slightly zygomorphic (in Oenothera), (3 or)4-merous;
stamens 2 times as many, or rarely as many, as sepals;
pollen usually shed in monads, rarely tetrads (Chylismia sect. Lignothera).
a dry capsule, usually dehiscent, sometimes indehiscent.
few to numerous, without hairs or wings, [very rarely with asymmetrical dry wing (Xylonagra)], or with dry (Oenothera), erose or smooth wing, or with thick, papillate wings (Chylismiella).
= 14, 28.
Oenothera hartwegii subsp. pubescens
Onagraceae tribe Onagreae
Subspecies pubescens occurs from Baca and Las Animas counties, Colorado, Clark, Meade, Morton, and Seward counties, Kansas, to western Oklahoma and the Texas Panhandle, throughout central and trans-Pecos Texas, west through eastern and southern New Mexico to central and southeastern Arizona, and also very locally in northern Mexico.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
Genera 13, species 265 (12 genera, 199 species in the flora).
Onagreae account for more than half the total genera in Onagraceae and diversified from a center in southwestern North America (L. Katinas et al. 2004). Delimitation of the tribe by W. L. Wagner et al. (2007) differs from previous ones by the exclusion of Gongylocarpus, now in its own tribe, by the segregation of eight genera (Camissoniopsis, Chylismia, Chylismiella, Eremothera, Eulobus, Neoholmgrenia, Taraxia, and Tetrapteron) from Camissonia, and by the inclusion of three previously separate genera (Calylophus, Gaura, and Stenosiphon) in Oenothera. Within the branch of the family that lacks stipules (Gongylocarpeae, Epilobieae, and Onagreae), the last two tribes form a clade that has very strong molecular support (R. A. Levin et al. 2003, 2004), but no obvious morphological synapomorphy. The clade may be defined by a cytogenetic change from the base chromosome number of x = 11 found in Circaeeae, Gongylocarpeae, and Lopezieae, to x = 18 in Epilobieae, and x = 7 in Onagreae; however, these changes could also have occurred independently. Other than the new chromosome number x = 7, the only apparent morphological synapomorphy for Onagreae alone is pollen with prominent apertural protrusions (J. Praglowski et al. 1987, 1989), a character state also found in Circaeeae (Praglowski et al. 1994). The monophyly of Onagreae has moderate (Levin et al. 2004) to strong support (V. S. Ford and L. D. Gottlieb 2007).
(Discussion copyrighted by Flora of North America; reprinted with permission.)
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