Oenothera harringtonii |
Oenothera heterophylla |
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Colorado Springs evening primrose |
variableleaf evening primrose |
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Habit | Herbs robust spring annual or, rarely, surviving a second year, caulescent, hirtellous, also glandular puberulent; from stout taproot. | Herbs annual or short-lived perennial, sparsely to densely strigillose, inflorescence sometimes also sparsely glandular puberulent, villous, or sparsely hirsute with spreading, pustulate-based hairs, or sometimes glabrate. | ||||
Stems | ascending to erect, stout, unbranched or with lateral stems from basal rosette, densely leafy, 15–30 cm. |
unbranched or branched mainly in distal part, 25–70 cm. |
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Leaves | 10–14(–14.5) × 1.5–2.3(–3) cm; petiole 4.3–6.6 cm; blade narrowly oblanceolate, margins irregularly and coarsely dentate, apex acute. |
in a basal rosette and cauline, basal 7–15 × 1–2.5 cm, cauline 3–13 ×0.4–2.3 cm; blade narrowly oblanceolate to oblanceolate, gradually narrowly lanceolate to lanceolate or elliptic distally, margins deeply lobed to remotely dentate or subentire; bracts longer than capsule they subtend, 1–3 cm. |
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Inflorescences | dense, often with several lateral branches, mature buds usually overtopping spike apex. |
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Flowers | usually 5–10 per stem opening per day near sunset, with heavy, sweet scent; buds erect; floral tube 31–60 mm; sepals 17–26 mm; petals white, fading pale pink, 20–26 mm; filaments 11–16 mm, anthers 8–11 mm; style 65–96 mm, stigma exserted beyond anthers at anthesis. |
2–several per spike opening per day near sunset; buds erect, with free tips erect or spreading, 1–6 mm; floral tube nearly straight, 25–47 mm; sepals 15–30 mm; petals yellow, broadly elliptic to nearly rhombic, 18–35 mm; filaments 15–30 mm, anthers 3–8 mm, pollen 85–100% fertile; style 45–75 mm, stigma usually exserted beyond anthers at anthesis. |
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Capsules | straight, lanceoloid, obtusely 4-angled, (21–)25–30(–35) × (5–)6–8 mm, tapering to a sterile beak 6–8 mm, dehiscent 1/2–2/3 their length, valve margins with 5–8 conspicuous, irregular tubercles, sometimes 2 or more coalesced into a sinuate ridge, also with conspicuous medial ridge throughout; pedicel 0.5–1 mm. |
lanceoloid, 13–25 × 2.5–4 mm. |
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Seeds | numerous, usually in 2 distinct rows per locule, sometimes rows partially overlapping, narrowly obovoid, 2.1–2.3 × 1–1.3 mm, embryo slightly less than 1/2 seed volume, surface appearing finely striate but papillose under magnification; seed collar with membrane intact at maturity, membrane rarely splitting and separating from collar, margin entire. |
brown, often flecked with darker spots, ellipsoid to broadly ellipsoid, 1.1–1.8 × 0.4–0.8 mm. |
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2n | = 14. |
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Oenothera harringtonii |
Oenothera heterophylla |
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Phenology | Flowering May–Jun. | |||||
Habitat | On compacted, silty clay to looser rocky and sandy soil in open grassland. | |||||
Elevation | 1400–1900 m. (4600–6200 ft.) | |||||
Distribution |
CO |
s United States |
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Discussion | Oenothera harringtonii is known only from southeastern Colorado from western El Paso and eastern Fremont counties, southeast through Pueblo to Otero counties, and south to Las Animas County; it may also occur in adjacent Colfax and Union counties in New Mexico but has not been collected there. Oenothera harringtonii is self-incompatible (W. L. Wagner et al. 1985; Wagner 2005). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Subspecies 2 (2 in the flora). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 10. | FNA vol. 10. | ||||
Parent taxa | ||||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | Raimannia heterophylla | |||||
Name authority | W. L. Wagner, Stockhouse & W. M. Klein: Ann. Missouri Bot. Gard. 70: 195. (1983) | Spach: Nouv. Ann. Mus. Hist. Nat. 4: 348. (1836) | ||||
Web links |