Oenothera grandis |
Oenothera cespitosa |
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largeflower eveningprimrose, showy evening-primrose |
fragrant evening-primrose, tuft evening-primrose |
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Habit | Herbs annual, strigillose and sparsely villous, also glandular puberulent distally. | Herbs perennial, acaulescent or caulescent, usually hirsute or villous, usually also glandular puberulent, or exclusively strigillose, rarely glabrous; from stout taproot, sometimes lateral roots producing adventitious shoots. | ||||||||||||||||||||||||||||
Stems | erect to ascending, often with ascending lateral branches, 15–60(–100) cm. |
(when present), usually ascending or decumbent, unbranched or branched from near base, 0–40 cm. |
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Leaves | in a basal rosette and cauline, basal 5–13 × 1–3 cm, cauline 3–10 × 1.5–3.5 cm; blade green, narrowly oblanceolate to narrowly elliptic, margins lobed or dentate, lobes often dentate; bracts spreading, flat. |
1.7–26(–36) × (0.3–)0.5–4.5(–6.5) cm; petiole (0.2–)1.7–11(–14) cm; blade usually oblanceolate to rhombic or spatulate, rarely elliptic, obovate, lanceolate, or linear-oblanceolate, margins irregularly sinuate-dentate, serrate, pinnatifid, lobed, or subentire, apex usually acute to rounded, rarely acuminate. |
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Flowers | 1–few opening per day near sunset; buds erect, with free tips terminal, erect or hornlike, 1.5–5 mm; floral tube 25–45 mm; sepals 15–30 mm; petals yellow, very broadly obovate or shallowly obcordate, 25–40 mm; filaments 12–22 mm, anthers 4–11 mm, pollen 85–100% fertile; style 40–75 mm, stigma exserted beyond anthers at anthesis. |
1–4(–6) per stem opening per day near sunset, with moderate to strong sweet scent with a rubbery background scent; buds usually erect, rarely recurved (during early development); floral tube (20–)40–140(–165) mm; sepals (15–)18–45(–54) mm; petals white, fading rose or rose pink to dark or deep rose purple, or pink to pale or light rose, or lavender, obovate or obcordate, (16–)20–50(–60) mm; filaments (6–)10–30(–35) mm, anthers (6–)9–17(–20) mm; style (45–)60–180(–185) mm, stigma exserted beyond anthers at anthesis. |
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Capsules | cylindrical, sometimes slightly enlarged toward apex, 25–50 ×2–3 mm. |
straight, curved, falcate, or sigmoid, usually cylindrical to lanceoloid or ellipsoid, sometimes ovoid, usually obtusely 4-angled, (10–)13–50(–68) × 4–9 mm, tapering to a sterile beak 6–8 mm, valve margins with rows of distinct tubercles to sinuate or nearly smooth ridges, dehiscent 1/3–7/8 their length; pedicel (0–)1–40(–55) mm. |
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Seeds | broadly ellipsoid to subglobose, 0.8–1.5 × 0.5–0.9 mm. |
numerous in 1 or 2 rows per locule, usually obovoid, oblong, or triangular, rarely suborbicular, 2.1–3.9 × 1–2.6 mm, embryo 1/5–2/3 of seed volume, surface papillose, reticulate or rarely irregularly roughened; seed collar sealed by a thin membrane, this flat or depressed into raphial cavity, when depressed often splitting, becoming separated from seed collar. |
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2n | = 14. |
= 14, 28. |
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Oenothera grandis |
Oenothera cespitosa |
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Phenology | Flowering Mar–Sep. | |||||||||||||||||||||||||||||
Habitat | Open, sandy sites. | |||||||||||||||||||||||||||||
Elevation | 0–1500(–2200) m. (0–4900(–7200) ft.) | |||||||||||||||||||||||||||||
Distribution |
AL; AR; CO; CT; FL; IL; IN; KS; LA; MD; MI; MO; NC; NE; NJ; NM; TX; Mexico (Tamaulipas)
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w North America; nw Mexico
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Discussion | Oenothera grandis is probably native to eastern New Mexico and Colorado, Arkansas, Kansas, Louisiana, Missouri, Nebraska, Oklahoma, and Texas, and northeastern Tamaulipas, Mexico. Scattered collections made in other states probably represent introductions (W. Dietrich and W. L. Wagner 1988). Oenothera grandis is self-incompatible (W. Dietrich and W. L. Wagner 1988). Oenothera laciniata Hill var. occidentalis Small and O. laciniata var. grandis Britton are illegitimate superfluous names based on O. sinuata Linnaeus var. grandiflora S. Watson and pertain here. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Subspecies 5 (5 in the flora). Oenothera cespitosa occurs in a wide array of habitats, from grassland, desert scrub, pinyon-juniper woodland, or Arizona chaparral to montane conifer forests, rarely at timberline, at elevations from (450–)800–3370 m. Oenothera cespitosa is self-incompatible (W. L. Wagner et al. 1985; Wagner 2005). Pachylophus nuttallii Spach is an illegitimate name that pertains here. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 10. | FNA vol. 10. | ||||||||||||||||||||||||||||
Parent taxa | Onagraceae > subfam. Onagroideae > tribe Onagreae > Oenothera > sect. Oenothera > subsect. Raimannia | Onagraceae > subfam. Onagroideae > tribe Onagreae > Oenothera > sect. Pachylophus | ||||||||||||||||||||||||||||
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Synonyms | O. laciniata var. grandiflora, O. sinuata var. grandiflora, Raimannia grandis | Pachylophus cespitosus | ||||||||||||||||||||||||||||
Name authority | Smyth: Trans. Kansas Acad. Sci. 16: 160. (1899) | Nuttall: Cat. Pl. Upper Louisiana, no. 53. (1813) | ||||||||||||||||||||||||||||
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