Oenothera glaucifolia
Onagraceae tribe Onagreae
false gaura
erect, branched or unbranched, 30–300 cm.
in a basal rosette and cauline, basal 3–7 × 0.5–2 cm, sessile, blade oblong to oblong-lanceolate, base usually ± auriculate, margins entire;
cauline 3–8(–10) × 0.4–1.8 cm, blade lanceolate to oblong-lanceolate, gradually smaller, becoming linear-subulate distally.
alternate or basal;
stipules absent.
long, wandlike, unbranched or branched.
4-merous, nearly actinomorphic, opening near sunrise;
floral tube 6–17 mm;
sepals 4–6 mm;
petals white, fading off-white or tinged pink, slightly unequal, rhombic, 4–6 mm, abruptly clawed;
filaments 5–8 mm, anthers 1.5–2 mm, pollen 85–100% fertile;
style 6–10, stigma exserted beyond anthers at anthesis.
usually actinomorphic, rarely slightly zygomorphic (in Oenothera), (3 or)4-merous;
stamens 2 times as many, or rarely as many, as sepals;
pollen usually shed in monads, rarely tetrads (Chylismia sect. Lignothera).
a dry capsule, usually dehiscent, sometimes indehiscent.
ovoid, 4-angled, somewhat flattened, 3–4 × 1.5–2.3 mm, valves with raised midrib and conspicuous lateral veins;
sessile.
1, pale yellow, oblanceoloid, 2.4–2.6 × 1–1.5 mm.
few to numerous, without hairs or wings, [very rarely with asymmetrical dry wing (Xylonagra)], or with dry (Oenothera), erose or smooth wing, or with thick, papillate wings (Chylismiella).
= 14.
Oenothera glaucifolia
Onagraceae tribe Onagreae
Oenothera glaucifolia is self-incompatible, the flowers diurnal, pollinated primarily by wasps (R. Clinebell, unpubl.), as well as bees, flies, butterflies, and occasionally beetles (summarized by W. L. Wagner et al. 2007). It was collected once in 1988 in Indiana at Miller Woods Visitor Center (Lake County), Dritz 596 (MOR); it seems likely that it was introduced, and has not been collected there since.
Stenosiphon virgatus Spach is a superfluous name and pertains here.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
Genera 13, species 265 (12 genera, 199 species in the flora).
Onagreae account for more than half the total genera in Onagraceae and diversified from a center in southwestern North America (L. Katinas et al. 2004). Delimitation of the tribe by W. L. Wagner et al. (2007) differs from previous ones by the exclusion of Gongylocarpus, now in its own tribe, by the segregation of eight genera (Camissoniopsis, Chylismia, Chylismiella, Eremothera, Eulobus, Neoholmgrenia, Taraxia, and Tetrapteron) from Camissonia, and by the inclusion of three previously separate genera (Calylophus, Gaura, and Stenosiphon) in Oenothera. Within the branch of the family that lacks stipules (Gongylocarpeae, Epilobieae, and Onagreae), the last two tribes form a clade that has very strong molecular support (R. A. Levin et al. 2003, 2004), but no obvious morphological synapomorphy. The clade may be defined by a cytogenetic change from the base chromosome number of x = 11 found in Circaeeae, Gongylocarpeae, and Lopezieae, to x = 18 in Epilobieae, and x = 7 in Onagreae; however, these changes could also have occurred independently. Other than the new chromosome number x = 7, the only apparent morphological synapomorphy for Onagreae alone is pollen with prominent apertural protrusions (J. Praglowski et al. 1987, 1989), a character state also found in Circaeeae (Praglowski et al. 1994). The monophyly of Onagreae has moderate (Levin et al. 2004) to strong support (V. S. Ford and L. D. Gottlieb 2007).
(Discussion copyrighted by Flora of North America; reprinted with permission.)
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