Oenothera glaucifolia |
Oenothera psammophila |
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false gaura |
St. Anthony dunes evening primrose |
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Habit | Herbs probably biennial, glabrous, becoming sparsely to densely glandular puberulent and short-villous distally, glaucous at least in proximal parts; from stout roots. | Herbs perennial, caulescent, glabrous, also with resinous exudate, especially on younger leaves; from woody taproot. |
Stems | erect, branched or unbranched, 30–300 cm. |
decumbent, usually branched, 10–30 cm, becoming woody and buried in sand. |
Leaves | in a basal rosette and cauline, basal 3–7 × 0.5–2 cm, sessile, blade oblong to oblong-lanceolate, base usually ± auriculate, margins entire; cauline 3–8(–10) × 0.4–1.8 cm, blade lanceolate to oblong-lanceolate, gradually smaller, becoming linear-subulate distally. |
(6–)8–9(–14.2) × (0.7–)1.5–2(–3.1) cm; petiole 3–9 cm; blade narrowly oblanceolate to oblanceolate, margins usually subentire or repand or remotely dentate, rarely serrate, apex acute. |
Inflorescences | long, wandlike, unbranched or branched. |
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Flowers | 4-merous, nearly actinomorphic, opening near sunrise; floral tube 6–17 mm; sepals 4–6 mm; petals white, fading off-white or tinged pink, slightly unequal, rhombic, 4–6 mm, abruptly clawed; filaments 5–8 mm, anthers 1.5–2 mm, pollen 85–100% fertile; style 6–10, stigma exserted beyond anthers at anthesis. |
1–3 per stem opening per day near sunset, with sweet scent; buds erect; floral tube 42–60 mm; sepals 22–28 mm; petals white, fading rose pink to rose, 23–40 mm; filaments 17–19 mm, anthers 13–16 mm; style 60–72(–88) mm, stigma exserted beyond anthers at anthesis. |
Capsules | ovoid, 4-angled, somewhat flattened, 3–4 × 1.5–2.3 mm, valves with raised midrib and conspicuous lateral veins; sessile. |
somewhat curved and often somewhat twisted, lanceoloid-cylindrical, nearly cylindrical, (20–)30–47 × 7–8 mm, gradually tapering to apex, 6–8 mm, dehiscent nearly throughout their length, valve margins with a conspicuous, irregular, wavy ridge; pedicel 1–5 mm. |
Seeds | 1, pale yellow, oblanceoloid, 2.4–2.6 × 1–1.5 mm. |
numerous, in 2 distinct rows per locule, narrowly obovoid, 2.5–3 × 1.2–1.4 mm, embryo 7/8 of seed volume, surface finely reticulate; seed collar with a broad membrane sealing cavity, margin entire. |
2n | = 14. |
= 14. |
Oenothera glaucifolia |
Oenothera psammophila |
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Phenology | Flowering May–Oct(–Nov). | Flowering May–Jul. |
Habitat | Rocky prairie slopes and outcrops or bluffs, along streams, roadsides, usuallyon limestone. | On barren areas of drifting sand at interface between outcrops of lava and sand dunes. |
Elevation | 200–1300 m. (700–4300 ft.) | 1500–1700 m. (4900–5600 ft.) |
Distribution |
AR; CO; KS; MO; NE; NM; OK; TX; WY
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ID |
Discussion | Oenothera glaucifolia is self-incompatible, the flowers diurnal, pollinated primarily by wasps (R. Clinebell, unpubl.), as well as bees, flies, butterflies, and occasionally beetles (summarized by W. L. Wagner et al. 2007). It was collected once in 1988 in Indiana at Miller Woods Visitor Center (Lake County), Dritz 596 (MOR); it seems likely that it was introduced, and has not been collected there since. Stenosiphon virgatus Spach is a superfluous name and pertains here. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Oenothera psammophila is known only from the dune area north and west of St. Anthony, Fremont County. It is unique in Oenothera because of the exudate produced on the leaves to which sand particles adhere, forming a sand sheath, presumably for protection from the constantly blowing sand particles. It is self-incompatible (W. L. Wagner et al. 1985; Wagner 2005). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 10. | FNA vol. 10. |
Parent taxa | Onagraceae > subfam. Onagroideae > tribe Onagreae > Oenothera > sect. Gaura > subsect. Stenosiphon | Onagraceae > subfam. Onagroideae > tribe Onagreae > Oenothera > sect. Pachylophus |
Sibling taxa | ||
Synonyms | Gaura linifolia, Stenosiphon linifolius | Pachylophus psammophilusa., O. cespitosa var. psammophila |
Name authority | W. L. Wagner & Hoch: Syst. Bot. Monogr. 83: 212. (2007) | (A. Nelson & J. F. Macbride) W. L. Wagner, Stockhouse & W. M. Klein: Monogr. Syst. Bot. Missouri Bot. Gard. 12: 84. (1985) |
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