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false gaura

Havard's evening primrose

Habit Herbs probably biennial, glabrous, becoming sparsely to densely glandular puberulent and short-villous distally, glaucous at least in proximal parts; from stout roots. Herbs compact to sprawling, strigillose; from a taproot, lateral roots producing adventitious shoots.
Stems

erect, branched or unbranched, 30–300 cm.

usually many-branched, sometimes unbranched, often twining among vegetation, sometimes rooting at nodes, 5–25(–70) cm.

Leaves

in a basal rosette and cauline, basal 3–7 × 0.5–2 cm, sessile, blade oblong to oblong-lanceolate, base usually ± auriculate, margins entire;

cauline 3–8(–10) × 0.4–1.8 cm, blade lanceolate to oblong-lanceolate, gradually smaller, becoming linear-subulate distally.

in a basal rosette and cauline, basal usually quickly deciduous, (1–)2–5 × (0.2–)0.5–1.5 cm;

petiole 0–0.6 cm;

blade oblanceolate, linear-lanceolate to linear distally, margins few toothed to pinnately lobed to sinuate-dentate distally.

Inflorescences

long, wandlike, unbranched or branched.

Flowers

4-merous, nearly actinomorphic, opening near sunrise;

floral tube 6–17 mm;

sepals 4–6 mm;

petals white, fading off-white or tinged pink, slightly unequal, rhombic, 4–6 mm, abruptly clawed;

filaments 5–8 mm, anthers 1.5–2 mm, pollen 85–100% fertile;

style 6–10, stigma exserted beyond anthers at anthesis.

1–few opening per day near sunset;

buds often twisted, free tips coherent;

floral tube (37–)45–60(–65) mm;

sepals (16–)18–26(–30) mm;

petals lemon-yellow, fading orange-red to reddish purple, usually elliptic, sometimes oblanceolate, (18–)21–30(–32) mm;

filaments 15–18(–22) mm, anthers red, 6–13 mm;

style (55–)65–86(–94) mm, stigma exserted beyond anthers at anthesis.

Capsules

ovoid, 4-angled, somewhat flattened, 3–4 × 1.5–2.3 mm, valves with raised midrib and conspicuous lateral veins;

sessile.

woody, narrowly ovoid to ovoid, 4-angled, 8–13(–16) × 3–4 mm, apex tapering to a short sterile beak 2–3 mm, valves with a prominent, broad midrib and capsule appearing 8-ribbed, tardily dehiscent ca. 1/3 capsule length.

Seeds

1, pale yellow, oblanceoloid, 2.4–2.6 × 1–1.5 mm.

2–2.5(–3.3) × 1.2–1.5 mm, sometimes with a small wing at distal end or a raised ridge along one longitudinal margin.

2n

= 14.

= 14, 28.

Oenothera glaucifolia

Oenothera havardii

Phenology Flowering May–Oct(–Nov). Flowering Apr–Oct.
Habitat Rocky prairie slopes and outcrops or bluffs, along streams, roadsides, usuallyon limestone. In depressions, seasonally wet flats, stream banks, margins of irrigated fields, sandy or clay soil, among tufted grasses like Sporobolus wrightii, primarily in Chihuahuan Desert.
Elevation 200–1300 m. (700–4300 ft.) 1300–2000 m. (4300–6600 ft.)
Distribution
from FNA
AR; CO; KS; MO; NE; NM; OK; TX; WY
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; TX; Mexico (Chihuahua, Durango, Sonora, Zacatecas)
[BONAP county map]
Discussion

Oenothera glaucifolia is self-incompatible, the flowers diurnal, pollinated primarily by wasps (R. Clinebell, unpubl.), as well as bees, flies, butterflies, and occasionally beetles (summarized by W. L. Wagner et al. 2007). It was collected once in 1988 in Indiana at Miller Woods Visitor Center (Lake County), Dritz 596 (MOR); it seems likely that it was introduced, and has not been collected there since.

Stenosiphon virgatus Spach is a superfluous name and pertains here.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Oenothera havardii ranges from Brewster and Presidio counties, Texas, and Cochise County, Arizona, south to Durango and Zacatecas, Mexico. W. L. Wagner (1984) found that O. havardii is self-incompatible and vespertine.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 10. FNA vol. 10.
Parent taxa Onagraceae > subfam. Onagroideae > tribe Onagreae > Oenothera > sect. Gaura > subsect. Stenosiphon Onagraceae > subfam. Onagroideae > tribe Onagreae > Oenothera > sect. Paradoxus
Sibling taxa
O. acutissima, O. albicaulis, O. argillicola, O. arida, O. arizonica, O. biennis, O. boquillensis, O. brachycarpa, O. calcicola, O. californica, O. canescens, O. capillifolia, O. cavernae, O. cespitosa, O. cinerea, O. clelandii, O. coloradensis, O. cordata, O. coronopifolia, O. coryi, O. curtiflora, O. curtissii, O. deltoides, O. demareei, O. dodgeniana, O. drummondii, O. elata, O. engelmannii, O. falfurriae, O. filiformis, O. filipes, O. flava, O. fruticosa, O. gaura, O. gayleana, O. glazioviana, O. grandiflora, O. grandis, O. harringtonii, O. hartwegii, O. havardii, O. heterophylla, O. hispida, O. howardii, O. humifusa, O. jamesii, O. kunthiana, O. laciniata, O. lavandulifolia, O. lindheimeri, O. linifolia, O. longissima, O. macrocarpa, O. mckelveyae, O. mexicana, O. nealleyi, O. neomexicana, O. nutans, O. nuttallii, O. oakesiana, O. organensis, O. pallida, O. parviflora, O. patriciae, O. perennis, O. pilosella, O. platanorum, O. podocarpa, O. primiveris, O. psammophila, O. pubescens, O. rhombipetala, O. riparia, O. rosea, O. serrulata, O. sessilis, O. simulans, O. sinuosa, O. spachiana, O. speciosa, O. stricta, O. suffrutescens, O. suffulta, O. tetraptera, O. texensis, O. toumeyi, O. triangulata, O. triloba, O. tubicula, O. villosa, O. wolfii, O. xylocarpa
O. acutissima, O. albicaulis, O. argillicola, O. arida, O. arizonica, O. biennis, O. boquillensis, O. brachycarpa, O. calcicola, O. californica, O. canescens, O. capillifolia, O. cavernae, O. cespitosa, O. cinerea, O. clelandii, O. coloradensis, O. cordata, O. coronopifolia, O. coryi, O. curtiflora, O. curtissii, O. deltoides, O. demareei, O. dodgeniana, O. drummondii, O. elata, O. engelmannii, O. falfurriae, O. filiformis, O. filipes, O. flava, O. fruticosa, O. gaura, O. gayleana, O. glaucifolia, O. glazioviana, O. grandiflora, O. grandis, O. harringtonii, O. hartwegii, O. heterophylla, O. hispida, O. howardii, O. humifusa, O. jamesii, O. kunthiana, O. laciniata, O. lavandulifolia, O. lindheimeri, O. linifolia, O. longissima, O. macrocarpa, O. mckelveyae, O. mexicana, O. nealleyi, O. neomexicana, O. nutans, O. nuttallii, O. oakesiana, O. organensis, O. pallida, O. parviflora, O. patriciae, O. perennis, O. pilosella, O. platanorum, O. podocarpa, O. primiveris, O. psammophila, O. pubescens, O. rhombipetala, O. riparia, O. rosea, O. serrulata, O. sessilis, O. simulans, O. sinuosa, O. spachiana, O. speciosa, O. stricta, O. suffrutescens, O. suffulta, O. tetraptera, O. texensis, O. toumeyi, O. triangulata, O. triloba, O. tubicula, O. villosa, O. wolfii, O. xylocarpa
Synonyms Gaura linifolia, Stenosiphon linifolius Hartmannia havardii, H. palmeri
Name authority W. L. Wagner & Hoch: Syst. Bot. Monogr. 83: 212. (2007) S. Watson: Proc. Amer. Acad. Arts 20: 366. (1885) — (as havardi)
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