Oenothera fruticosa
Onagraceae subfam. onagroideae
narrow-leaf evening-primrose
erect to decumbent, branched or unbranched, (10–)30–80 cm.
in a basal rosette and cauline, rosette usually withered by anthesis, surfaces sometimes glaucous, especially abaxially, basal 3–12 × 0.5–3 cm, petiole 1–4 cm, blade oblanceolate to obovate, margins subentire, dentate, or denticulate, sometimes undulate;
cauline 2–6(–11) × (0.1–)0.5–2(–5) cm, petiole 0.1–2(–6) cm, blade linear, lanceolate to oblanceolate, narrowly elliptic, or ovate, margins subentire or dentate or denticulate, sometimes undulate.
stipules present or absent.
usually erect, rarely nodding, flowers in axils of distalmost few nodes.
opening near sunrise;
buds with free tips 0.5–8(–13) mm, connivent, sometimes spreading;
floral tube 5–20 mm;
sepals 5–20 mm;
petals pale to bright yellow, fading pale pink, orangish pink, or yellow, (8–)15–25(–30) mm;
filaments 5–15 mm, anthers 4–7 mm, pollen 85–100% fertile;
style 12–20 mm, stigma exserted beyond anthers at anthesis.
floral tube present or, rarely, absent;
sepals 2 or 4 (very rarely 3), deciduous with floral tube, petals, and stamens;
petals yellow, white, pink, red, rarely in combination.
clavate to oblong-clavate or oblong-ellipsoid, 4-angled to 4-winged, (5–)10–17(–20) × (2–)3–4(–6) mm, stipe 0.1–10 mm;
sessile.
1 × 0.5 mm.
Oenothera fruticosa
Onagraceae subfam. onagroideae
Subspecies 2 (2 in the flora).
Oenothera fruticosa as delimited here is a polymorphic species. Previous classification of this group has undergone numerous reorganizations due to the difficulties in separating populations into discrete morphological taxa. In the past it has most frequently been treated as two species, O. fruticosa and O. tetragona Roth, often with a dozen or more infraspecific taxa recognized. The broad delimitation of G. B. Straley (1977) is followed here with one species consisting of two subspecies that appear to intergrade extensively across a wide area of overlap. Straley determined O. fruticosa to be self-incompatible and polyploid.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
Genera 21, species 582 (16 genera, 246 species in the flora).
Onagroideae encompass the main lineage of the family, after the early branching of Ludwigia (R. A. Levin et al. 2003, 2004). This large and diverse lineage is distinguished by the presence of a floral tube beyond the apex of the ovary; sepals deciduous with the floral tube, petals, and stamens; pollen shed in monads (or tetrads in Chylismia sect. Lignothera and all but one species of Epilobium); ovular vascular system exclusively transseptal (R. H. Eyde 1981); ovule archesporium multicellular (H. Tobe and P. H. Raven 1996); and change in base chromosome number from x = 8 in Ludwigia to x = 10 or x = 11 at the base of Onagroideae (Raven 1979; Levin et al. 2003). Molecular work (Levin et al. 2003, 2004) substantially supports the traditional tribal classification (P. A. Munz 1965; Raven 1979, 1988); tribes are recognized to delimit major branches within the phylogeny of Onagroideae, where the branches comprise strongly supported monophyletic groups of one or more genera.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
1. Capsules clavate to oblong-clavate, widest above middle; plants moderately to densely strigillose, and/or sometimes glandular puberulent or villous; leaf blade margins subentire or weakly and remotely denticulate. | subsp. fruticosa |
1. Capsules oblong to oblong-ellipsoid, widest at middle; plants glandular puberulent or glabrous, sometimes villous or strigillose; leaf blade margins dentate to remotely denticulate. | subsp. tetragona |
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