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beach evening-primrose

Habit Herbs annual, sparsely villous proximally, leaves glabrate to sparsely villous along veins and on margins, usually glandular puberulent in distal parts; from stout taproot.
Stems

usually well-branched, 20–70(–100) cm.

Leaves

in a basal rosette and cauline, basal 3.5–9 × 0.5–1.5 cm, blade lyrate;

cauline 1.5–7 × 0.1–0.6 cm, blade narrowly lanceolate to linear, margins sinuate-dentate, undulate.

Flowers

4-merous, zygomorphic, opening at sunset;

floral tube 10–20 mm;

sepals 11–21 mm;

petals white, fading pink to red, elliptic to elliptic-obovate, 10–15 mm;

filaments 8–13 mm, anthers 2–6 mm, pollen 90–100% fertile;

style 22–36 mm, stigma exserted beyond anthers at anthesis.

Capsules

ellipsoid or ovoid, narrowly 4-winged, furrowed between angles, 4.5–8 × 2–5 mm, stipe 0.2–2.2 mm;

sessile.

Seeds

3 or 4 (or 5), yellowish to light brown, 2–3(–4) × 1 mm.

Subspecies

thalassaphila (Brandegee) W.

Dietrich

& W.

l

.

Wagner

differs from subsp. drummondii in a number of modally distinctive morphological features, especially floral tubes 2–3.5 cm, sepal tips 0.3–1 mm, capsules 2–4 cm × 2.5–5 mm in diameter and those, coupled with the great disjunction from the Atlantic coast of the United States and Mexico to the southern tip of Baja California, make it worthy of recognition.

Oenothera

drummondii is self-compatible and outcrossing.

2n

= 14.

Oenothera drummondii

Oenothera nealleyi

Phenology Flowering Apr–Oct.
Habitat Washes, sandy places, grass­lands, extending to pinyon-juniper woodlands.
Elevation 1200–2200 m. (3900–7200 ft.)
Distribution
from USDA
n Mexico; s United States
[WildflowerSearch map]
[BONAP county map]
from FNA
NM; TX; Mexico (Coahuila)
Discussion

Subspecies 2 (1 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Oenothera nealleyi is restricted to an area from trans-Pecos Texas and northern Coahuila, Mexico, north to Bernalillo and Torrance counties, New Mexico. P. H. Raven and D. P. Gregory (1972[1973]) considered O. nealleyi to represent an unevenly intergrading entity with O. suffulta based on merging of distinguishing characteristics. The known intermediates occur in Terrell County, Texas, and were previously described as Gaura suffulta var. terrellensis Munz, but until new data on its status are available, we include this name with O. nealleyi. The molecular data (K. N. Krakos, unpubl.) suggest that O. nealleyi is not as closely related to O. suffulta as suggested by Raven and Gregory, given the placement in the phylogeny and the difference in scent profiles for these two taxa. Oenothera suffulta is a member of a strongly supported clade that also includes O. patriciae and O. triangulata, while O. nealleyi is a member of a polytomy that consists of other species of subsect. Gaura, with the O. suffultaO. triangulataO. patriciae clade sister to it (W. L. Wagner et al. 2013). Oenothera nealleyi has a strong sweet scent, whereas O. suffulta does not have a discernible scent (Wagner et al.). Raven and Gregory determined O. nealleyi to be self-incompatible.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 10. FNA vol. 10.
Parent taxa Onagraceae > subfam. Onagroideae > tribe Onagreae > Oenothera > sect. Oenothera > subsect. Raimannia Onagraceae > subfam. Onagroideae > tribe Onagreae > Oenothera > sect. Gaura > subsect. Gaura
Sibling taxa
O. acutissima, O. albicaulis, O. argillicola, O. arida, O. arizonica, O. biennis, O. boquillensis, O. brachycarpa, O. calcicola, O. californica, O. canescens, O. capillifolia, O. cavernae, O. cespitosa, O. cinerea, O. clelandii, O. coloradensis, O. cordata, O. coronopifolia, O. coryi, O. curtiflora, O. curtissii, O. deltoides, O. demareei, O. dodgeniana, O. elata, O. engelmannii, O. falfurriae, O. filiformis, O. filipes, O. flava, O. fruticosa, O. gaura, O. gayleana, O. glaucifolia, O. glazioviana, O. grandiflora, O. grandis, O. harringtonii, O. hartwegii, O. havardii, O. heterophylla, O. hispida, O. howardii, O. humifusa, O. jamesii, O. kunthiana, O. laciniata, O. lavandulifolia, O. lindheimeri, O. linifolia, O. longissima, O. macrocarpa, O. mckelveyae, O. mexicana, O. nealleyi, O. neomexicana, O. nutans, O. nuttallii, O. oakesiana, O. organensis, O. pallida, O. parviflora, O. patriciae, O. perennis, O. pilosella, O. platanorum, O. podocarpa, O. primiveris, O. psammophila, O. pubescens, O. rhombipetala, O. riparia, O. rosea, O. serrulata, O. sessilis, O. simulans, O. sinuosa, O. spachiana, O. speciosa, O. stricta, O. suffrutescens, O. suffulta, O. tetraptera, O. texensis, O. toumeyi, O. triangulata, O. triloba, O. tubicula, O. villosa, O. wolfii, O. xylocarpa
O. acutissima, O. albicaulis, O. argillicola, O. arida, O. arizonica, O. biennis, O. boquillensis, O. brachycarpa, O. calcicola, O. californica, O. canescens, O. capillifolia, O. cavernae, O. cespitosa, O. cinerea, O. clelandii, O. coloradensis, O. cordata, O. coronopifolia, O. coryi, O. curtiflora, O. curtissii, O. deltoides, O. demareei, O. dodgeniana, O. drummondii, O. elata, O. engelmannii, O. falfurriae, O. filiformis, O. filipes, O. flava, O. fruticosa, O. gaura, O. gayleana, O. glaucifolia, O. glazioviana, O. grandiflora, O. grandis, O. harringtonii, O. hartwegii, O. havardii, O. heterophylla, O. hispida, O. howardii, O. humifusa, O. jamesii, O. kunthiana, O. laciniata, O. lavandulifolia, O. lindheimeri, O. linifolia, O. longissima, O. macrocarpa, O. mckelveyae, O. mexicana, O. neomexicana, O. nutans, O. nuttallii, O. oakesiana, O. organensis, O. pallida, O. parviflora, O. patriciae, O. perennis, O. pilosella, O. platanorum, O. podocarpa, O. primiveris, O. psammophila, O. pubescens, O. rhombipetala, O. riparia, O. rosea, O. serrulata, O. sessilis, O. simulans, O. sinuosa, O. spachiana, O. speciosa, O. stricta, O. suffrutescens, O. suffulta, O. tetraptera, O. texensis, O. toumeyi, O. triangulata, O. triloba, O. tubicula, O. villosa, O. wolfii, O. xylocarpa
Subordinate taxa
O. drummondii subsp. drummondii
Synonyms Raimannia drummondii Gaura nealleyi, G. suffulta subsp. nealleyi, G. suffulta var. terrellensis, O. suffulta subsp. nealleyi
Name authority Hooker: Bot. Mag. 61: plate 3361. (1834) (J. M. Coulter) Krakos & W. L. Wagner: PhytoKeys 28: 64. (2013)
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