Oenothera dodgeniana
Onagraceae tribe Onagreae
1 or few-branched from base, 50–120 cm.
in a basal rosette and cauline, basal 6–20 × 1–3 cm, blade lanceolate to narrowly elliptic;
cauline 5–10 × 1–2.5 cm, blade lanceolate to narrowly elliptic, margins subentire or repand-denticulate.
alternate or basal;
stipules absent.
4-merous, zygomorphic, opening at sunset;
floral tube 10–11 mm;
sepals 11–15 mm;
petals white, fading pink, rhombic-obovate, 11–14 mm;
filaments 6.5–9 mm, anthers 2.5–4 mm, pollen 90–100% fertile;
style 22–28 mm, stigma exserted beyond anthers.
usually actinomorphic, rarely slightly zygomorphic (in Oenothera), (3 or)4-merous;
stamens 2 times as many, or rarely as many, as sepals;
pollen usually shed in monads, rarely tetrads (Chylismia sect. Lignothera).
a dry capsule, usually dehiscent, sometimes indehiscent.
ellipsoid or ovoid, sharply 4-angled, with deep furrows alternating with angles for 2–3 mm from apex, ribbed from base of furrow to base of fruit, 9–11 × 3–5 mm;
sessile.
2–4, yellowish to light brown, 2–3 mm.
few to numerous, without hairs or wings, [very rarely with asymmetrical dry wing (Xylonagra)], or with dry (Oenothera), erose or smooth wing, or with thick, papillate wings (Chylismiella).
= 14.
Oenothera dodgeniana
Onagraceae tribe Onagreae
Oenothera dodgeniana occurs in two disjunct areas: the western foothills of the San Juan Mountains in Archuleta and Huerfano counties, Colorado, and Rio Arriba County, New Mexico; and Sierra Blanca and Sacramento Mountains in Lincoln and Otero counties, south-central New Mexico. The species was collected once at Durango, La Plata County, Colorado (P. H. Raven and D. P. Gregory 1972[1973]), but has not since been recollected. Oenothera dodgeniana and O. coloradensis were considered by Raven and Gregory to represent a relict species along the eastern flank of the Rocky Mountains that arose from more widespread species farther to the east, such as O. filiformis. Oenothera dodgeniana belongs to a subclade which is sister to that containing O. coloradensis, and within that subclade is sister to O. demareei and O. lindheimeri (W. L. Wagner et al. 2013). Although O. dodgeniana is fairly closely related to O. coloradensis, the two taxa seem to have had independent origins that have led to distributions along the eastern flank of the Rocky Mountains. Oenothera dodgeniana is self-compatible (Raven and Gregory).
(Discussion copyrighted by Flora of North America; reprinted with permission.)
Genera 13, species 265 (12 genera, 199 species in the flora).
Onagreae account for more than half the total genera in Onagraceae and diversified from a center in southwestern North America (L. Katinas et al. 2004). Delimitation of the tribe by W. L. Wagner et al. (2007) differs from previous ones by the exclusion of Gongylocarpus, now in its own tribe, by the segregation of eight genera (Camissoniopsis, Chylismia, Chylismiella, Eremothera, Eulobus, Neoholmgrenia, Taraxia, and Tetrapteron) from Camissonia, and by the inclusion of three previously separate genera (Calylophus, Gaura, and Stenosiphon) in Oenothera. Within the branch of the family that lacks stipules (Gongylocarpeae, Epilobieae, and Onagreae), the last two tribes form a clade that has very strong molecular support (R. A. Levin et al. 2003, 2004), but no obvious morphological synapomorphy. The clade may be defined by a cytogenetic change from the base chromosome number of x = 11 found in Circaeeae, Gongylocarpeae, and Lopezieae, to x = 18 in Epilobieae, and x = 7 in Onagreae; however, these changes could also have occurred independently. Other than the new chromosome number x = 7, the only apparent morphological synapomorphy for Onagreae alone is pollen with prominent apertural protrusions (J. Praglowski et al. 1987, 1989), a character state also found in Circaeeae (Praglowski et al. 1994). The monophyly of Onagreae has moderate (Levin et al. 2004) to strong support (V. S. Ford and L. D. Gottlieb 2007).
(Discussion copyrighted by Flora of North America; reprinted with permission.)
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