Oenothera dodgeniana
Oenothera nealleyi
1 or few-branched from base, 50–120 cm.
usually well-branched, 20–70(–100) cm.
in a basal rosette and cauline, basal 6–20 × 1–3 cm, blade lanceolate to narrowly elliptic;
cauline 5–10 × 1–2.5 cm, blade lanceolate to narrowly elliptic, margins subentire or repand-denticulate.
in a basal rosette and cauline, basal 3.5–9 × 0.5–1.5 cm, blade lyrate;
cauline 1.5–7 × 0.1–0.6 cm, blade narrowly lanceolate to linear, margins sinuate-dentate, undulate.
4-merous, zygomorphic, opening at sunset;
floral tube 10–11 mm;
sepals 11–15 mm;
petals white, fading pink, rhombic-obovate, 11–14 mm;
filaments 6.5–9 mm, anthers 2.5–4 mm, pollen 90–100% fertile;
style 22–28 mm, stigma exserted beyond anthers.
4-merous, zygomorphic, opening at sunset;
floral tube 10–20 mm;
sepals 11–21 mm;
petals white, fading pink to red, elliptic to elliptic-obovate, 10–15 mm;
filaments 8–13 mm, anthers 2–6 mm, pollen 90–100% fertile;
style 22–36 mm, stigma exserted beyond anthers at anthesis.
ellipsoid or ovoid, sharply 4-angled, with deep furrows alternating with angles for 2–3 mm from apex, ribbed from base of furrow to base of fruit, 9–11 × 3–5 mm;
sessile.
ellipsoid or ovoid, narrowly 4-winged, furrowed between angles, 4.5–8 × 2–5 mm, stipe 0.2–2.2 mm;
sessile.
2–4, yellowish to light brown, 2–3 mm.
3 or 4 (or 5), yellowish to light brown, 2–3(–4) × 1 mm.
= 14.
= 14.
Oenothera dodgeniana
Oenothera nealleyi
Oenothera dodgeniana occurs in two disjunct areas: the western foothills of the San Juan Mountains in Archuleta and Huerfano counties, Colorado, and Rio Arriba County, New Mexico; and Sierra Blanca and Sacramento Mountains in Lincoln and Otero counties, south-central New Mexico. The species was collected once at Durango, La Plata County, Colorado (P. H. Raven and D. P. Gregory 1972[1973]), but has not since been recollected. Oenothera dodgeniana and O. coloradensis were considered by Raven and Gregory to represent a relict species along the eastern flank of the Rocky Mountains that arose from more widespread species farther to the east, such as O. filiformis. Oenothera dodgeniana belongs to a subclade which is sister to that containing O. coloradensis, and within that subclade is sister to O. demareei and O. lindheimeri (W. L. Wagner et al. 2013). Although O. dodgeniana is fairly closely related to O. coloradensis, the two taxa seem to have had independent origins that have led to distributions along the eastern flank of the Rocky Mountains. Oenothera dodgeniana is self-compatible (Raven and Gregory).
(Discussion copyrighted by Flora of North America; reprinted with permission.)
Oenothera nealleyi is restricted to an area from trans-Pecos Texas and northern Coahuila, Mexico, north to Bernalillo and Torrance counties, New Mexico. P. H. Raven and D. P. Gregory (1972[1973]) considered O. nealleyi to represent an unevenly intergrading entity with O. suffulta based on merging of distinguishing characteristics. The known intermediates occur in Terrell County, Texas, and were previously described as Gaura suffulta var. terrellensis Munz, but until new data on its status are available, we include this name with O. nealleyi. The molecular data (K. N. Krakos, unpubl.) suggest that O. nealleyi is not as closely related to O. suffulta as suggested by Raven and Gregory, given the placement in the phylogeny and the difference in scent profiles for these two taxa. Oenothera suffulta is a member of a strongly supported clade that also includes O. patriciae and O. triangulata, while O. nealleyi is a member of a polytomy that consists of other species of subsect. Gaura, with the O. suffulta—O. triangulata—O. patriciae clade sister to it (W. L. Wagner et al. 2013). Oenothera nealleyi has a strong sweet scent, whereas O. suffulta does not have a discernible scent (Wagner et al.). Raven and Gregory determined O. nealleyi to be self-incompatible.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
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