Oenothera demareei |
Oenothera argillicola |
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demaree's beeblossom |
shale barren evening primrose |
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Habit | Herbs usually robust winter-annual, sometimes biennial, densely strigillose throughout; from fleshy taproot. | Herbs biennial or short-lived perennial, strigillose and sparsely to moderately villous, hairs sometimes pustulate, pustules with green or red bases, inflorescence glabrous or sparsely glandular puberulent, sometimes also sparsely villous. |
Stems | usually well-branched distal to base, 50–400 cm. |
erect to ascending, green or red, unbranched or with branches obliquely arising from rosette or in distal 1/2 of main stem. |
Leaves | in a basal rosette and cauline, 3–7 × 0.2–0.8 cm; blade narrowly elliptic to elliptic or lanceolate, margins subentire or shallowly undulate-denticulate. |
in a basal rosette and cauline, basal 7–25 × 0.7–2 cm, cauline 6–13 × 0.4–1 cm; blade dark green, somewhat glossy, very narrowly oblanceolate to narrowly oblanceolate, linear-elliptic, lanceolate, or nearly linear, margins flat, entire or remotely and bluntly dentate, sometimes with larger teeth near base; bracts persistent. |
Inflorescences | curved with ascending tip, unbranched. |
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Flowers | 4-merous, zygomorphic, opening at sunrise; floral tube 4–13(–15) mm; sepals 13–20 mm; petals white, fading pink, rhombic-obovate, 10–17 mm; filaments 8–17 mm, anthers 3–7 mm, pollen 90–100% fertile; style 18–32 mm, stigma exserted beyond anthers at anthesis. |
opening near sunset; buds erect, 4–8 mm diam., with free tips subterminal, divergent and hornlike, 3–9 mm; floral tube 32–52 mm; sepals yellowish green to yellow, sometimes flushed with red, especially at apex, 27–38 mm; petals yellow to pale yellow, fading pale yellow to pale yellowish orange, very broadly obcordate or obovate, 25–42 mm; filaments 20–27 mm, anthers 9–13 mm, pollen 90–100% fertile; style 60–85 mm, stigma exserted beyond anthers at anthesis. |
Capsules | ellipsoid or ovoid, sharply 4-angled, 4.5–7 × 1.5–2.5 mm; sessile. |
spreading at nearly a right angle to stem, curved upward, sometimes secund, dull green or rusty brown when dry, narrowly lanceoloid to lanceoloid, 20–40 × 4–6 mm, free tips of valves 1–2 mm. |
Seeds | 2–4, yellowish to reddish brown, 1.2–3 × 0.7–1.3 mm. |
1.3–1.9 × 0.7–1.1 mm. |
2n | = 14. |
= 14. |
Oenothera demareei |
Oenothera argillicola |
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Phenology | Flowering Jul–Oct. | Flowering (Jun–)Jul–Oct. |
Habitat | Open meadows in sandy loam. | Open sites on Devonian Brallier shale slopes, barrens, outcrops or adjacent roadsides in mid-Appalachian Allegheny Mountains. |
Elevation | 70–200 m. (200–700 ft.) | 150–700 m. (500–2300 ft.) |
Distribution |
AR |
MD; PA; VA; WV |
Discussion | Oenothera demareei is known only from Clark, Garland, Hempstead, Howard, Montgomery, Pike, Saline, and Sevier counties. P. H. Raven and D. P. Gregory (1972[1973]) found Oenothera demareei to be self-incompatible. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Oenothera argillicola is one of eight angiosperm species restricted to the Devonian Brallier shale barrens, but among them only O. argillicola and Trifolium virginicum occur throughout the shale barren region. Oenothera argillicola has plastome V and a CC genome composition. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 10. | FNA vol. 10. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | Gaura demareei | O. argillicola var. pubescens |
Name authority | (P. H. Raven & D. P. Gregory) W. L. Wagner & Hoch: Syst. Bot. Monogr. 83: 212. (2007) | Mackenzie: Torreya 4: 56. (1904) |
Web links |