Oenothera deltoides
Onagraceae tribe Onagreae
basket evening-primrose, birdcage evening primrose, desert lantern, devil's lantern, dune primrose, hairy evening primrose, lion-in-a-cage
central stem usually erect, usually thickened at base and spongy, branched or unbranched, branches few–several, slender, decumbent to ascending, from base, usually encircling central stem in older plants, 10–40(–100) cm.
in a basal rosette and cauline, rosette usually well developed (except subsp. howellii), basal 5–25 × 1–5 cm, cauline 4–12(–18) × 0.5–4 cm;
petiole 1.5–8 cm;
blade rhombic-obovate, lanceolate, or oblanceolate, margins subentire, dentate, or pinnatifid.
alternate or basal;
stipules absent.
1–several opening per day near sunset;
buds nodding, weakly or strongly quadrangular or fluted in distal 1/2, with free tips 0–9 mm;
floral tube 20–40 mm;
sepals (13–)15–35 mm, not spotted;
petals white, fading pink to deep pink, broadly obovate or obcordate, 15–44 mm;
filaments 8–15 mm, anthers 5–14 mm;
style 35–60 mm, stigma exserted beyond anthers at anthesis.
usually actinomorphic, rarely slightly zygomorphic (in Oenothera), (3 or)4-merous;
stamens 2 times as many, or rarely as many, as sepals;
pollen usually shed in monads, rarely tetrads (Chylismia sect. Lignothera).
a dry capsule, usually dehiscent, sometimes indehiscent.
spreading, straight to curved, becoming somewhat woody in age, cylindrical to slightly 4-angled, widest toward base, tapering from base to apex, (15–)30–80 × 1.5–5 mm;
sessile.
numerous, in 1 row per locule, buff with dark spots or black, narrowly obovoid, 1.5–2.8 mm.
few to numerous, without hairs or wings, [very rarely with asymmetrical dry wing (Xylonagra)], or with dry (Oenothera), erose or smooth wing, or with thick, papillate wings (Chylismiella).
Oenothera deltoides
Onagraceae tribe Onagreae
Subspecies 5 (5 in the flora).
Oenothera deltoides is self-incompatible or self-compatible (W. M. Klein 1964; W. L. Wagner et al. 2007; K. E. Theiss et al. 2010).
(Discussion copyrighted by Flora of North America; reprinted with permission.)
Genera 13, species 265 (12 genera, 199 species in the flora).
Onagreae account for more than half the total genera in Onagraceae and diversified from a center in southwestern North America (L. Katinas et al. 2004). Delimitation of the tribe by W. L. Wagner et al. (2007) differs from previous ones by the exclusion of Gongylocarpus, now in its own tribe, by the segregation of eight genera (Camissoniopsis, Chylismia, Chylismiella, Eremothera, Eulobus, Neoholmgrenia, Taraxia, and Tetrapteron) from Camissonia, and by the inclusion of three previously separate genera (Calylophus, Gaura, and Stenosiphon) in Oenothera. Within the branch of the family that lacks stipules (Gongylocarpeae, Epilobieae, and Onagreae), the last two tribes form a clade that has very strong molecular support (R. A. Levin et al. 2003, 2004), but no obvious morphological synapomorphy. The clade may be defined by a cytogenetic change from the base chromosome number of x = 11 found in Circaeeae, Gongylocarpeae, and Lopezieae, to x = 18 in Epilobieae, and x = 7 in Onagreae; however, these changes could also have occurred independently. Other than the new chromosome number x = 7, the only apparent morphological synapomorphy for Onagreae alone is pollen with prominent apertural protrusions (J. Praglowski et al. 1987, 1989), a character state also found in Circaeeae (Praglowski et al. 1994). The monophyly of Onagreae has moderate (Levin et al. 2004) to strong support (V. S. Ford and L. D. Gottlieb 2007).
(Discussion copyrighted by Flora of North America; reprinted with permission.)
1. Herbs perennial, from long, fleshy roots; stems not thickened at base, branches erect or ascending, not encircling stems in older plants. | subsp. howellii |
1. Herbs annual or short-lived perennial from a taproot; stems usually thickened at base, branches ascending or decumbent, often encircling stems in older plants. | → 2 |
2. Flower buds fluted or strongly quadrangular in distal 1/2, without free tips, villous with curly hairs, sometimes glabrous; leaf blade margins sinuate-dentate to pinnatifid; petals 15–25(–30) mm; capsules 15–25(–30) mm. | subsp. piperi |
2. Flower buds weakly or strongly quadrangular in distal 1/2, with free tips 0–3 mm, strigillose, sparsely to moderately villous, or glabrous; leaf blade margins sinuate-dentate or subentire, rarely pinnatifid; petals 15–44 mm; capsules 40–80 mm. | → 3 |
3. Flower buds with free tips 1–3 mm, quadrangular in distal 1/2; plants strigillose, especially distally. | subsp. ambigua |
3. Flower buds with free tips 0–1.5 mm, weakly quadrangular in distal 1/2; plants glabrous, villous, or strigillose. | → 4 |
4. Herbs annual, strigillose, sometimes also villous; capsules 2–3.5 mm diam.; flower buds with free tips 0–1.5 mm. | subsp. deltoides |
4. Herbs short-lived perennial or sometimes annual, glabrous or sparsely villous, rarely also strigillose; capsules 3–5 mm diam.; flower buds without free tips. | subsp. cognata |
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OR
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