Oenothera deltoides |
Oenothera nealleyi |
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basket evening-primrose, birdcage evening primrose, desert lantern, devil's lantern, dune primrose, hairy evening primrose, lion-in-a-cage |
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Habit | Herbs usually winter-annual, sometimes perennial, glabrous, glandular puberulent, strigillose, and/or villous, sometimes more villous distally, hairs sometimes very curly, especially on flower parts; from a taproot or relatively long, fleshy roots. | Herbs annual, sparsely villous proximally, leaves glabrate to sparsely villous along veins and on margins, usually glandular puberulent in distal parts; from stout taproot. | ||||||||||||||||
Stems | central stem usually erect, usually thickened at base and spongy, branched or unbranched, branches few–several, slender, decumbent to ascending, from base, usually encircling central stem in older plants, 10–40(–100) cm. |
usually well-branched, 20–70(–100) cm. |
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Leaves | in a basal rosette and cauline, rosette usually well developed (except subsp. howellii), basal 5–25 × 1–5 cm, cauline 4–12(–18) × 0.5–4 cm; petiole 1.5–8 cm; blade rhombic-obovate, lanceolate, or oblanceolate, margins subentire, dentate, or pinnatifid. |
in a basal rosette and cauline, basal 3.5–9 × 0.5–1.5 cm, blade lyrate; cauline 1.5–7 × 0.1–0.6 cm, blade narrowly lanceolate to linear, margins sinuate-dentate, undulate. |
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Flowers | 1–several opening per day near sunset; buds nodding, weakly or strongly quadrangular or fluted in distal 1/2, with free tips 0–9 mm; floral tube 20–40 mm; sepals (13–)15–35 mm, not spotted; petals white, fading pink to deep pink, broadly obovate or obcordate, 15–44 mm; filaments 8–15 mm, anthers 5–14 mm; style 35–60 mm, stigma exserted beyond anthers at anthesis. |
4-merous, zygomorphic, opening at sunset; floral tube 10–20 mm; sepals 11–21 mm; petals white, fading pink to red, elliptic to elliptic-obovate, 10–15 mm; filaments 8–13 mm, anthers 2–6 mm, pollen 90–100% fertile; style 22–36 mm, stigma exserted beyond anthers at anthesis. |
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Capsules | spreading, straight to curved, becoming somewhat woody in age, cylindrical to slightly 4-angled, widest toward base, tapering from base to apex, (15–)30–80 × 1.5–5 mm; sessile. |
ellipsoid or ovoid, narrowly 4-winged, furrowed between angles, 4.5–8 × 2–5 mm, stipe 0.2–2.2 mm; sessile. |
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Seeds | numerous, in 1 row per locule, buff with dark spots or black, narrowly obovoid, 1.5–2.8 mm. |
3 or 4 (or 5), yellowish to light brown, 2–3(–4) × 1 mm. |
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2n | = 14. |
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Oenothera deltoides |
Oenothera nealleyi |
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Phenology | Flowering Apr–Oct. | |||||||||||||||||
Habitat | Washes, sandy places, grasslands, extending to pinyon-juniper woodlands. | |||||||||||||||||
Elevation | 1200–2200 m. (3900–7200 ft.) | |||||||||||||||||
Distribution |
w United States; nw Mexico
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NM; TX; Mexico (Coahuila) |
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Discussion | Subspecies 5 (5 in the flora). Oenothera deltoides is self-incompatible or self-compatible (W. M. Klein 1964; W. L. Wagner et al. 2007; K. E. Theiss et al. 2010). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Oenothera nealleyi is restricted to an area from trans-Pecos Texas and northern Coahuila, Mexico, north to Bernalillo and Torrance counties, New Mexico. P. H. Raven and D. P. Gregory (1972[1973]) considered O. nealleyi to represent an unevenly intergrading entity with O. suffulta based on merging of distinguishing characteristics. The known intermediates occur in Terrell County, Texas, and were previously described as Gaura suffulta var. terrellensis Munz, but until new data on its status are available, we include this name with O. nealleyi. The molecular data (K. N. Krakos, unpubl.) suggest that O. nealleyi is not as closely related to O. suffulta as suggested by Raven and Gregory, given the placement in the phylogeny and the difference in scent profiles for these two taxa. Oenothera suffulta is a member of a strongly supported clade that also includes O. patriciae and O. triangulata, while O. nealleyi is a member of a polytomy that consists of other species of subsect. Gaura, with the O. suffulta—O. triangulata—O. patriciae clade sister to it (W. L. Wagner et al. 2013). Oenothera nealleyi has a strong sweet scent, whereas O. suffulta does not have a discernible scent (Wagner et al.). Raven and Gregory determined O. nealleyi to be self-incompatible. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 10. | FNA vol. 10. | ||||||||||||||||
Parent taxa | Onagraceae > subfam. Onagroideae > tribe Onagreae > Oenothera > sect. Anogra | Onagraceae > subfam. Onagroideae > tribe Onagreae > Oenothera > sect. Gaura > subsect. Gaura | ||||||||||||||||
Sibling taxa | ||||||||||||||||||
Subordinate taxa | ||||||||||||||||||
Synonyms | Anogra deltoides | Gaura nealleyi, G. suffulta subsp. nealleyi, G. suffulta var. terrellensis, O. suffulta subsp. nealleyi | ||||||||||||||||
Name authority | Torrey & Frémont in J. C. Frémont: Rep. Exped. Rocky Mts., 315. (1845) | (J. M. Coulter) Krakos & W. L. Wagner: PhytoKeys 28: 64. (2013) | ||||||||||||||||
Web links |