Oenothera cespitosa subsp. macroglottis
Onagraceae tribe Onagreae
(if present), usually unbranched, rarely with 1–several short laterals, 4–8 cm.
(6.8–)9.5–23(–32) × (1.3–)2.4–4.5(–6.5) cm;
petiole (3–)4–11(–14) cm;
blade oblanceolate to spatulate, margins often undulate, usually regularly to irregularly dentate, rarely coarsely and irregularly pinnately lobed.
alternate or basal;
stipules absent.
floral tube (45–)75–110(–153) mm;
sepals (22–)30–45(–50) mm;
petals fading pink to pale rose, (21–)35–43(–50) mm;
filaments (16–)19–28(–35) mm, anthers (10–)12–17 mm;
style 85–180 mm.
usually actinomorphic, rarely slightly zygomorphic (in Oenothera), (3 or)4-merous;
stamens 2 times as many, or rarely as many, as sepals;
pollen usually shed in monads, rarely tetrads (Chylismia sect. Lignothera).
a dry capsule, usually dehiscent, sometimes indehiscent.
somewhat curved, lanceoloid-cylindrical to cylindrical, symmetrical throughout, sometimes slightly flattened on one side at base, (17–)25–45(–56) × 5–8 mm, valve margins with conspicuous, nearly smooth to irregular undulate ridge;
pedicel 2–7 mm.
narrowly obovoid, 2.5–3 × 1–1.4 mm, embryo 1/2 of seed volume, surface minutely papillose to reticulate;
seed collar forming narrow slit above raphe with a slightly sunken membrane, margin entire or obscurely sinuate distally.
few to numerous, without hairs or wings, [very rarely with asymmetrical dry wing (Xylonagra)], or with dry (Oenothera), erose or smooth wing, or with thick, papillate wings (Chylismiella).
= 14, 28.
Oenothera cespitosa subsp. macroglottis
Onagraceae tribe Onagreae
Genera 13, species 265 (12 genera, 199 species in the flora).
Onagreae account for more than half the total genera in Onagraceae and diversified from a center in southwestern North America (L. Katinas et al. 2004). Delimitation of the tribe by W. L. Wagner et al. (2007) differs from previous ones by the exclusion of Gongylocarpus, now in its own tribe, by the segregation of eight genera (Camissoniopsis, Chylismia, Chylismiella, Eremothera, Eulobus, Neoholmgrenia, Taraxia, and Tetrapteron) from Camissonia, and by the inclusion of three previously separate genera (Calylophus, Gaura, and Stenosiphon) in Oenothera. Within the branch of the family that lacks stipules (Gongylocarpeae, Epilobieae, and Onagreae), the last two tribes form a clade that has very strong molecular support (R. A. Levin et al. 2003, 2004), but no obvious morphological synapomorphy. The clade may be defined by a cytogenetic change from the base chromosome number of x = 11 found in Circaeeae, Gongylocarpeae, and Lopezieae, to x = 18 in Epilobieae, and x = 7 in Onagreae; however, these changes could also have occurred independently. Other than the new chromosome number x = 7, the only apparent morphological synapomorphy for Onagreae alone is pollen with prominent apertural protrusions (J. Praglowski et al. 1987, 1989), a character state also found in Circaeeae (Praglowski et al. 1994). The monophyly of Onagreae has moderate (Levin et al. 2004) to strong support (V. S. Ford and L. D. Gottlieb 2007).
(Discussion copyrighted by Flora of North America; reprinted with permission.)
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