Oenothera cespitosa subsp. macroglottis |
Onagraceae subfam. onagroideae |
|
---|---|---|
|
||
Habit | Herbs acaulescent or short-caulescent, hirsute and glandular puberulent, or glabrous. | |
Stems | (if present), usually unbranched, rarely with 1–several short laterals, 4–8 cm. |
|
Leaves | (6.8–)9.5–23(–32) × (1.3–)2.4–4.5(–6.5) cm; petiole (3–)4–11(–14) cm; blade oblanceolate to spatulate, margins often undulate, usually regularly to irregularly dentate, rarely coarsely and irregularly pinnately lobed. |
stipules present or absent. |
Flowers | floral tube (45–)75–110(–153) mm; sepals (22–)30–45(–50) mm; petals fading pink to pale rose, (21–)35–43(–50) mm; filaments (16–)19–28(–35) mm, anthers (10–)12–17 mm; style 85–180 mm. |
floral tube present or, rarely, absent; sepals 2 or 4 (very rarely 3), deciduous with floral tube, petals, and stamens; petals yellow, white, pink, red, rarely in combination. |
Capsules | somewhat curved, lanceoloid-cylindrical to cylindrical, symmetrical throughout, sometimes slightly flattened on one side at base, (17–)25–45(–56) × 5–8 mm, valve margins with conspicuous, nearly smooth to irregular undulate ridge; pedicel 2–7 mm. |
|
Seeds | narrowly obovoid, 2.5–3 × 1–1.4 mm, embryo 1/2 of seed volume, surface minutely papillose to reticulate; seed collar forming narrow slit above raphe with a slightly sunken membrane, margin entire or obscurely sinuate distally. |
|
x |
||
2n | = 14, 28. |
|
Oenothera cespitosa subsp. macroglottis |
Onagraceae subfam. onagroideae |
|
Phenology | Flowering May–Jul(–Sep). | |
Habitat | Open, igneous rocky slopes, talus, roadcuts, open or shaded and sandy or gravelly sites along streams, rarely on shale, in upper pinyon-juniper woodlands, Gambel oak scrub, ponderosa pine forests, ponderosa pine-Douglas fir forests, spruce-fir-lodgepole pine forests. | |
Elevation | 2000–3100 m. (6600–10200 ft.) | |
Distribution |
CO; NM; UT; WY |
North America; Mexico; Central America; South America; West Indies; Eurasia; Pacific Islands (New Zealand, Society Islands); Australia |
Discussion | Genera 21, species 582 (16 genera, 246 species in the flora). Onagroideae encompass the main lineage of the family, after the early branching of Ludwigia (R. A. Levin et al. 2003, 2004). This large and diverse lineage is distinguished by the presence of a floral tube beyond the apex of the ovary; sepals deciduous with the floral tube, petals, and stamens; pollen shed in monads (or tetrads in Chylismia sect. Lignothera and all but one species of Epilobium); ovular vascular system exclusively transseptal (R. H. Eyde 1981); ovule archesporium multicellular (H. Tobe and P. H. Raven 1996); and change in base chromosome number from x = 8 in Ludwigia to x = 10 or x = 11 at the base of Onagroideae (Raven 1979; Levin et al. 2003). Molecular work (Levin et al. 2003, 2004) substantially supports the traditional tribal classification (P. A. Munz 1965; Raven 1979, 1988); tribes are recognized to delimit major branches within the phylogeny of Onagroideae, where the branches comprise strongly supported monophyletic groups of one or more genera. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
|
Source | FNA vol. 10. | FNA vol. 10. |
Parent taxa | ||
Sibling taxa | ||
Subordinate taxa | ||
Synonyms | Pachylophus macroglottis, O. cespitosa var. macroglottis, P. hirsutus | |
Name authority | (Rydberg) W. L. Wagner, Stockhouse & W. M. Klein: Ann. Missouri Bot. Gard. 70: 195. (1983) — (as caespitosa) | W. L. Wagner & Hoch: Syst. Bot. Monogr. 83: 41. (2007) |
Web links |