Oenothera cespitosa subsp. macroglottis |
Oenothera cespitosa |
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fragrant evening-primrose, tuft evening-primrose |
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Habit | Herbs acaulescent or short-caulescent, hirsute and glandular puberulent, or glabrous. | Herbs perennial, acaulescent or caulescent, usually hirsute or villous, usually also glandular puberulent, or exclusively strigillose, rarely glabrous; from stout taproot, sometimes lateral roots producing adventitious shoots. | ||||||||||||||||||||||||||||
Stems | (if present), usually unbranched, rarely with 1–several short laterals, 4–8 cm. |
(when present), usually ascending or decumbent, unbranched or branched from near base, 0–40 cm. |
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Leaves | (6.8–)9.5–23(–32) × (1.3–)2.4–4.5(–6.5) cm; petiole (3–)4–11(–14) cm; blade oblanceolate to spatulate, margins often undulate, usually regularly to irregularly dentate, rarely coarsely and irregularly pinnately lobed. |
1.7–26(–36) × (0.3–)0.5–4.5(–6.5) cm; petiole (0.2–)1.7–11(–14) cm; blade usually oblanceolate to rhombic or spatulate, rarely elliptic, obovate, lanceolate, or linear-oblanceolate, margins irregularly sinuate-dentate, serrate, pinnatifid, lobed, or subentire, apex usually acute to rounded, rarely acuminate. |
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Flowers | floral tube (45–)75–110(–153) mm; sepals (22–)30–45(–50) mm; petals fading pink to pale rose, (21–)35–43(–50) mm; filaments (16–)19–28(–35) mm, anthers (10–)12–17 mm; style 85–180 mm. |
1–4(–6) per stem opening per day near sunset, with moderate to strong sweet scent with a rubbery background scent; buds usually erect, rarely recurved (during early development); floral tube (20–)40–140(–165) mm; sepals (15–)18–45(–54) mm; petals white, fading rose or rose pink to dark or deep rose purple, or pink to pale or light rose, or lavender, obovate or obcordate, (16–)20–50(–60) mm; filaments (6–)10–30(–35) mm, anthers (6–)9–17(–20) mm; style (45–)60–180(–185) mm, stigma exserted beyond anthers at anthesis. |
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Capsules | somewhat curved, lanceoloid-cylindrical to cylindrical, symmetrical throughout, sometimes slightly flattened on one side at base, (17–)25–45(–56) × 5–8 mm, valve margins with conspicuous, nearly smooth to irregular undulate ridge; pedicel 2–7 mm. |
straight, curved, falcate, or sigmoid, usually cylindrical to lanceoloid or ellipsoid, sometimes ovoid, usually obtusely 4-angled, (10–)13–50(–68) × 4–9 mm, tapering to a sterile beak 6–8 mm, valve margins with rows of distinct tubercles to sinuate or nearly smooth ridges, dehiscent 1/3–7/8 their length; pedicel (0–)1–40(–55) mm. |
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Seeds | narrowly obovoid, 2.5–3 × 1–1.4 mm, embryo 1/2 of seed volume, surface minutely papillose to reticulate; seed collar forming narrow slit above raphe with a slightly sunken membrane, margin entire or obscurely sinuate distally. |
numerous in 1 or 2 rows per locule, usually obovoid, oblong, or triangular, rarely suborbicular, 2.1–3.9 × 1–2.6 mm, embryo 1/5–2/3 of seed volume, surface papillose, reticulate or rarely irregularly roughened; seed collar sealed by a thin membrane, this flat or depressed into raphial cavity, when depressed often splitting, becoming separated from seed collar. |
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2n | = 14, 28. |
= 14, 28. |
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Oenothera cespitosa subsp. macroglottis |
Oenothera cespitosa |
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Phenology | Flowering May–Jul(–Sep). | |||||||||||||||||||||||||||||
Habitat | Open, igneous rocky slopes, talus, roadcuts, open or shaded and sandy or gravelly sites along streams, rarely on shale, in upper pinyon-juniper woodlands, Gambel oak scrub, ponderosa pine forests, ponderosa pine-Douglas fir forests, spruce-fir-lodgepole pine forests. | |||||||||||||||||||||||||||||
Elevation | 2000–3100 m. (6600–10200 ft.) | |||||||||||||||||||||||||||||
Distribution |
CO; NM; UT; WY |
w North America; nw Mexico
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Discussion | Subspecies 5 (5 in the flora). Oenothera cespitosa occurs in a wide array of habitats, from grassland, desert scrub, pinyon-juniper woodland, or Arizona chaparral to montane conifer forests, rarely at timberline, at elevations from (450–)800–3370 m. Oenothera cespitosa is self-incompatible (W. L. Wagner et al. 1985; Wagner 2005). Pachylophus nuttallii Spach is an illegitimate name that pertains here. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 10. | FNA vol. 10. | ||||||||||||||||||||||||||||
Parent taxa | ||||||||||||||||||||||||||||||
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Synonyms | Pachylophus macroglottis, O. cespitosa var. macroglottis, P. hirsutus | Pachylophus cespitosus | ||||||||||||||||||||||||||||
Name authority | (Rydberg) W. L. Wagner, Stockhouse & W. M. Klein: Ann. Missouri Bot. Gard. 70: 195. (1983) — (as caespitosa) | Nuttall: Cat. Pl. Upper Louisiana, no. 53. (1813) | ||||||||||||||||||||||||||||
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