Oenothera cespitosa subsp. crinita |
Oenothera cespitosa |
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cespitose evening-primrose |
fragrant evening-primrose, tuft evening-primrose |
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Habit | Herbs acaulescent or caulescent, densely hirsute, also sparsely glandular puberulent. | Herbs perennial, acaulescent or caulescent, usually hirsute or villous, usually also glandular puberulent, or exclusively strigillose, rarely glabrous; from stout taproot, sometimes lateral roots producing adventitious shoots. | ||||||||||||||||||||||||||||
Stems | unbranched to many-branched, and then sometimes producing dense clumps 5–50 cm diam., 2–14 cm. |
(when present), usually ascending or decumbent, unbranched or branched from near base, 0–40 cm. |
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Leaves | 1.7–10(–18) × (0.3–)0.5–2.5(–3.4) cm; petiole (0.2–)3–5(–8) cm; blade usually oblanceolate to linear-oblanceolate, rarely obovate, margins subentire, sinuate, or dentate to pinnatifid. |
1.7–26(–36) × (0.3–)0.5–4.5(–6.5) cm; petiole (0.2–)1.7–11(–14) cm; blade usually oblanceolate to rhombic or spatulate, rarely elliptic, obovate, lanceolate, or linear-oblanceolate, margins irregularly sinuate-dentate, serrate, pinnatifid, lobed, or subentire, apex usually acute to rounded, rarely acuminate. |
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Flowers | floral tube (28–)35–75(–85) mm; sepals (15–)18–25(–27) mm; petals fading rose or sometimes deep rose purple, (16–)20–30(–35) mm; filaments (6–)10–17(–20) mm, anthers 6–8(–10) mm; style (45–)60–90(–105) mm. |
1–4(–6) per stem opening per day near sunset, with moderate to strong sweet scent with a rubbery background scent; buds usually erect, rarely recurved (during early development); floral tube (20–)40–140(–165) mm; sepals (15–)18–45(–54) mm; petals white, fading rose or rose pink to dark or deep rose purple, or pink to pale or light rose, or lavender, obovate or obcordate, (16–)20–50(–60) mm; filaments (6–)10–30(–35) mm, anthers (6–)9–17(–20) mm; style (45–)60–180(–185) mm, stigma exserted beyond anthers at anthesis. |
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Capsules | usually falcate or sigmoid, especially when young, also somewhat flattened, ellipsoid-ovoid to lanceoloid, 10–31(–34) × 4–9 mm, valve margins with 8–15 tubercles or these coalesced into a sinuate ridge; pedicel 0.5–1 mm. |
straight, curved, falcate, or sigmoid, usually cylindrical to lanceoloid or ellipsoid, sometimes ovoid, usually obtusely 4-angled, (10–)13–50(–68) × 4–9 mm, tapering to a sterile beak 6–8 mm, valve margins with rows of distinct tubercles to sinuate or nearly smooth ridges, dehiscent 1/3–7/8 their length; pedicel (0–)1–40(–55) mm. |
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Seeds | obovoid, oblong, or ± triangular, 2.9–3.5 × 1.1–2 mm, embryo 1/2–2/3 of seed volume, surface papillose, reticulate or very minutely roughened; seed collar membrane depressed and often splitting, becoming separated from collar at maturity, margin conspicuously sinuate throughout, surface often ribbed, ribs forming partial or complete vertical partitions in collar. |
numerous in 1 or 2 rows per locule, usually obovoid, oblong, or triangular, rarely suborbicular, 2.1–3.9 × 1–2.6 mm, embryo 1/5–2/3 of seed volume, surface papillose, reticulate or rarely irregularly roughened; seed collar sealed by a thin membrane, this flat or depressed into raphial cavity, when depressed often splitting, becoming separated from seed collar. |
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2n | = 14, 28. |
= 14, 28. |
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Oenothera cespitosa subsp. crinita |
Oenothera cespitosa |
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Phenology | Flowering Apr–Jul. | |||||||||||||||||||||||||||||
Habitat | Open sites, compacted or loose soil derived from dolomite, limestone, tufa, or marble, exposed knolls, gravelly benches, steep slopes, scree, rocky mesas, rocky arroyos, from mountain summits in alpine or subalpine communities with Pinus longaeva and P. flexilis or pinyon-juniper woodlands to Great Basin or Mojave Desert shrub communities dominated by Artemisia, Atriplex confertifolia, Coleogyne, Hilaria, Lycium. | |||||||||||||||||||||||||||||
Elevation | 1100–3400 m. (3600–11200 ft.) | |||||||||||||||||||||||||||||
Distribution |
AZ; CA; NV; UT |
w North America; nw Mexico
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Discussion | Subspecies crinita is the most polymorphic subspecies of Oenothera cespitosa; it is also the least understood. W. L. Wagner et al. (1985) grouped two series of populations that appear to intergrade together within the limits of this subspecies. One population is a morphologically relatively uniform form characterized by a many-branched habit, which may form dense clumps to 50 cm diameter, leaves that are 2–7 cm, floral tubes 25–60 mm and petals that fade to a rose color, and it occurs at high elevations on rocky, limestone sites or at lower elevations on extreme, chalky, white limestone and dolomite substrates or sometimes scree slopes. A more common form occurs at low to mid elevations in pinyon-juniper woodlands to Great Basin or Mojave Desert scrub on rocky slopes, talus, or along arroyos that is much less compact with one to several clustered rosettes, rarely more, with leaves 8–16 cm, floral tubes 45–75 mm, and petals that fade rose purple. The common form also grows on limestone and dolomite but, unlike the clumped form, it does not seem to be restricted to it. To compound the problem, many foothill and valley populations of subsp. crinita intergrade extensively with subspp. cespitosa and marginata. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Subspecies 5 (5 in the flora). Oenothera cespitosa occurs in a wide array of habitats, from grassland, desert scrub, pinyon-juniper woodland, or Arizona chaparral to montane conifer forests, rarely at timberline, at elevations from (450–)800–3370 m. Oenothera cespitosa is self-incompatible (W. L. Wagner et al. 1985; Wagner 2005). Pachylophus nuttallii Spach is an illegitimate name that pertains here. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 10. | FNA vol. 10. | ||||||||||||||||||||||||||||
Parent taxa | ||||||||||||||||||||||||||||||
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Synonyms | Pachylophus crinitus, O. cespitosa var. crinita, O. cespitosa subsp. jonesii, O. cespitosa var. jonesii, O. cespitosa var. stellae | Pachylophus cespitosus | ||||||||||||||||||||||||||||
Name authority | (Rydberg) Munz in N. L. Britton et al.: N. Amer. Fl., ser. 2, 5: 100. (1965) — (as caespitosa) | Nuttall: Cat. Pl. Upper Louisiana, no. 53. (1813) | ||||||||||||||||||||||||||||
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