Oenothera capillifolia |
Oenothera primiveris |
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desert evening primrose, yellow desert evening primrose |
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Habit | Herbs perennial (short-lived or, sometimes, suffrutescent) or annual, glabrous or strigillose; from a stout taproot. | Herbs winter-annual, caulescent to short-caulescent, long-hirsute, hairs often with reddish purple pustulate bases, especially proximally, also moderately strigillose, and glandular puberulent distally, often on leaves; from a weakly fleshy taproot. | ||||
Stems | 1–many, weakly decumbent to ascending or erect, unbranched to moderately branched, (10–)25–80 cm. |
(when present) unbranched and erect or, sometimes, few branches from near base, in robust plants stems and caudex hollow and greatly enlarged, especially toward base, densely leafy, 5–35 cm. |
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Leaves | 1–9 × (0.1–)0.3–1 cm, sometimes fascicles of small leaves to 2 cm present in non-flowering axils; petiole 0–0.6 cm; blade linear to narrowly lanceolate or oblanceolate, often folded lengthwise, usually not much reduced distally, proximalmost leaves sometimes spatulate, base attenuate, margins subentire or serrulate or spinulose-serrate, apex acute. |
in a basal rosette, sometimes also cauline, (1.4–)6–15(–28) × (0.2–)1–3.5(–5.6) cm; petiole (0.9–)3.5–8(–14) cm; blade oblanceolate to linear-oblanceolate, pinnatifid or 2-pinnatifid to shallowly pinnately lobed, margins sinuate-dentate or subentire, apex obtuse. |
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Flowers | opening at sunrise; buds with free tips 0–4 mm; floral tube 5–20 mm; sepals 4–12 mm, midribs keeled; petals yellow, fading orangish to purplish, 6–25 mm; antisepalous filaments 2–8 mm, antipetalous filaments 1–4 mm, anthers 2–7 mm, pollen 90–100% fertile; style 9–30 mm, stigma sometimes blue-black, discoid to quadrangular, exserted beyond anthers. |
usually 1–4, rarely more, opening per day, 1–2 hours before sunset; sepals (7–)12–25(–30) mm; petals yellow, fading reddish orange to purple, obcordate to obovate, (6–)13–35(–40) mm; filaments 6–16 mm, anthers 3–10 mm; style (32–)40–90(–100) mm, stigma exserted beyond anthers or surrounded by them. |
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Capsules | 10–35 × 1–2 mm, hard, dehiscent 1/2 their length, often tardily dehiscent throughout their length. |
woody in age, sigmoid or curved to nearly straight, lanceoloid to ovoid, 4-angled, 10–45(–60) × 4–8 mm, beak 4–15 mm, dehiscent 1/4–2/3 their length; sessile. |
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Seeds | obovoid, 1–1.8 mm, sharply angled, apex truncate. |
usually numerous, in 2 rows per locule, obovoid to oblanceoloid, 3–3.5 × 1–1.4 mm, surface thickened above raphe and at distal end into U-shaped structure. |
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2n | = 14. |
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Oenothera capillifolia |
Oenothera primiveris |
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Phenology | Flowering Feb–May(–Jun). | |||||
Habitat | Sandy soil on flats, low hills and margins of sand dunes, along arroyos, roadsides, in desert scrub, grasslands and oak-grasslands. | |||||
Elevation | 30–1600 m. (100–5200 ft.) | |||||
Distribution | c United States; sc United States; n Mexico |
AZ; CA; NM; NV; TX; UT; Mexico (Baja California, Chihuahua, Sonora)
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Discussion | Subspecies 2 (2 in the flora). Oenothera capillifolia is self-incompatible (H. F. Towner 1977). Oenothera berlandieri (Spach) Steudel 1841, not D. Dietrich 1840, is superfluous and cannot be used in Oenothera when transferred from Calylophus, and pertains here. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Oenothera primiveris has a complex variation pattern (W. L. Wagner 2005). In the western part of the range from southeastern California across southern Nevada to southern Utah counties of Emery, Kane, and Washington, and northwestern Mohave County, Arizona, plants generally have a gray appearance, with dense pubescence and larger flowers with widespread self-compatibility, but with scattered populations retaining self-incompatibility. Populations from southof the Mogollon Plateau to southern New Mexico, western Texas, Chihuahua, Sonora, and Baja California, Mexico, are greener in appearance with smaller to much smaller flowers, and are all self-compatible with occasional outcrossing or complete autogamy. The transitions between these two extremes are so extensive and more or less gradual that it is not possible to subdivide into two subspecies as has been done previously (Wagner). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 10. | FNA vol. 10. | ||||
Parent taxa | Onagraceae > subfam. Onagroideae > tribe Onagreae > Oenothera > sect. Calylophus > subsect. Calylophus | Onagraceae > subfam. Onagroideae > tribe Onagreae > Oenothera > sect. Eremia | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | Meriolix capillifolia | Lavauxia lobata, L. primiveris, O. bufonis, O. cespitosa var. primiveris, O. johnsonii, O. primiveris subsp. bufonis, O. primiveris var. bufonis, O. primiveris subsp. caulescens, O. primiveris var. caulescens, Pachylophus johnsonii | ||||
Name authority | Scheele: Linnaea 21: 576. (1848) | A. Gray: Smithsonian Contr. Knowl. 5(6): 58. (1853) | ||||
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