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Eureka dunes evening primrose

Habit Herbs perennial, densely strigillose and villous; with deep-seated, fleshy underground parts from underground horizontal rootstocks. Herbs (annual or perennial), [shrubs].
Stems

sprawling to decumbent, sometimes new rosettes forming at stem apex when becoming buried in drifting sand, 15–60 cm.

Leaves

blade rhombic-ovate to oblanceolate, margins entire or weakly dentate.

alternate or basal;

stipules absent.

Flowers

floral tube 25–30 mm;

sepals 15–25 mm;

petals 20–30 mm.

usually actinomorphic, rarely slightly zygomorphic (in Oenothera), (3 or)4-merous;

stamens 2 times as many, or rarely as many, as sepals;

pollen usually shed in monads, rarely tetrads (Chylismia sect. Lignothera).

Fruit

a dry capsule, usually dehiscent, sometimes indehiscent.

Capsules

30–70 mm.

Seeds

few to numerous, without hairs or wings, [very rarely with asymmetrical dry wing (Xylonagra)], or with dry (Oenothera), erose or smooth wing, or with thick, papillate wings (Chylismiella).

2n

= 14.

Oenothera californica subsp. eurekensis

Onagraceae tribe Onagreae

Phenology Flowering (Mar)Jun–Jul(Sep).
Habitat Sand dunes.
Elevation 900–1200 m. (3000–3900 ft.)
Distribution
from FNA
CA
[BONAP county map]
North America; Mexico; Central America; South America; West Indies
Discussion

Subspecies eurekensis is known from three main areas within the Eureka Dunes system, Inyo County. It is federally listed as endangered and is in the Center for Plant Conservation’s National Collection of Endangered Plants.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Genera 13, species 265 (12 genera, 199 species in the flora).

Onagreae account for more than half the total genera in Onagraceae and diversified from a center in southwestern North America (L. Katinas et al. 2004). Delimitation of the tribe by W. L. Wagner et al. (2007) differs from previous ones by the exclusion of Gongylocarpus, now in its own tribe, by the segregation of eight genera (Camissoniopsis, Chylismia, Chylismiella, Eremothera, Eulobus, Neoholmgrenia, Taraxia, and Tetrapteron) from Camissonia, and by the inclusion of three previously separate genera (Calylophus, Gaura, and Stenosiphon) in Oenothera. Within the branch of the family that lacks stipules (Gongylocarpeae, Epilobieae, and Onagreae), the last two tribes form a clade that has very strong molecular support (R. A. Levin et al. 2003, 2004), but no obvious morphological synapomorphy. The clade may be defined by a cytogenetic change from the base chromosome number of x = 11 found in Circaeeae, Gongylocarpeae, and Lopezieae, to x = 18 in Epilobieae, and x = 7 in Onagreae; however, these changes could also have occurred independently. Other than the new chromosome number x = 7, the only apparent morphological synapomorphy for Onagreae alone is pollen with prominent apertural protrusions (J. Praglowski et al. 1987, 1989), a character state also found in Circaeeae (Praglowski et al. 1994). The monophyly of Onagreae has moderate (Levin et al. 2004) to strong support (V. S. Ford and L. D. Gottlieb 2007).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 10. FNA vol. 10.
Parent taxa Onagraceae > subfam. Onagroideae > tribe Onagreae > Oenothera > sect. Anogra > Oenothera californica Onagraceae > subfam. Onagroideae
Sibling taxa
O. californica subsp. avita, O. californica subsp. californica
Subordinate taxa
Synonyms O. deltoides subsp. eurekensis, O. avita subsp. eurekensis
Name authority (Munz & J. C. Roos) W. M. Klein: Aliso 5: 179. (1962) Dumortier: Fl. Belg., 89. (1827)
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