Oenothera calcicola |
Oenothera nutans |
|
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Texas beeblossom |
nodding evening-primrose |
|
Habit | Herbs perennial, clumped, usually sparsely strigillose, rarely glabrate or sparsely villous, hairs erect, sometimes sparsely glandular puberulent distally; from twisted, woody rootstock. | Herbs biennial, often appearing glabrous to naked eye, usually strigillose and sparsely to moderately villous proximal to inflorescence, hairs translucent and with or without pustules, pustules not red (in fresh material), inflorescence glabrous, glandular puberulent, or strigillose and glandular puberulent. |
Stems | erect, branched below and just above ground, branched also proximal to inflorescences, (40–)60–250 cm. |
erect, red on proximal parts, green on distal ones, rarely red throughout, unbranched or with branches obliquely arising from rosette and secondary branches arising from main stem, 30–200 cm. |
Leaves | in a basal rosette and cauline, basal 3–13 × 0.6–2.5 cm, blade spatulate to oblanceolate, cauline (1–)2.5–12 × 0.1–1 cm, blade linear to narrowly oblanceolate, margins slightly to conspicuously sinuate-dentate. |
in a basal rosette and cauline, basal 10–32 × 3–7 cm, cauline 6–20 × 2–8 cm; blade green to pale green, narrowly oblanceolate to narrowly obovate, or narrowly elliptic, sometimes lanceolate distally, margins usually flat, rarely undulate, bluntly dentate, teeth widely spaced, sometimes sinuate-dentate proximally; bracts caducous. |
Inflorescences | slender. |
erect, unbranched or with secondary branches just proximal to main one. |
Flowers | 4-merous, zygomorphic, opening near sunset; floral tube 3–9 mm; sepals 6–12 mm; petals white, fading dark pink to red, slightly unequal, elliptic-obovate, 7–11 mm; stamens presented in lower 1/2 of flower, filaments 3–7 mm, anthers 2.5–5 mm, pollen 90–100% fertile; style 9.5–19 mm, stigma exserted beyond anthers at anthesis. |
opening near sunset; buds erect, 4–6 mm diam., with free tips terminal, erect, 1.5–6 mm; floral tube 30–43 mm; sepals yellowish green or flushed with red, 10–23 mm; petals yellow to pale yellow, fading pale yellowish white, very broadly obcordate, 14–25(–30) mm; filaments 10–25 mm, anthers 4–10 mm, pollen ca. 50% fertile; style 35–63 mm, stigma surrounded by anthers at anthesis. |
Capsules | lanceoloid to narrowly ovoid, narrowly 4-winged, 7–12 × 1.5–2.5 mm, tapered to a sterile stipe 2–5 mm. |
erect or slightly spreading, dull green when dry, narrowly lanceoloid to narrowly ovoid, 12–36 × 3–6 mm, free tips of valves 1–1.5 mm. |
Seeds | (2 or)3 or 4(or 5), light brown or reddish brown, 1.5–2.5 × 0.8–1.3 mm. |
1–1.9 × 0.6–0.9 mm. |
2n | = 14. |
= 14. |
Oenothera calcicola |
Oenothera nutans |
|
Phenology | Flowering Apr–May. | Flowering Jun–Aug(–Sep). |
Habitat | Dry limestone, gypsum, or caliche soil, slopes. | Open, often disturbed sites, stream beds, flood plains, slopes, margins of mixed deciduous forests, roadsides, old fields. |
Elevation | 400–1800(–2100) m. (1300–5900(–6900) ft.) | (200–)400–1700 m. ((700–)1300–5600 ft.) |
Distribution |
TX; Mexico (Coahuila, Durango, Nuevo León, Tamaulipas) |
AL; AR; CT; DE; FL; GA; IN; KY; MA; MD; ME; MI; MO; NC; NH; NJ; NY; OH; PA; SC; TN; VA; VT; WV; ON |
Discussion | Oenothera calcicola occurs at mostly higher elevations and more montane areas than other species of subsect. Stipogaura, from the southern Trans-Pecos and Edwards Plateau regions of Texas southward into northern Mexico. P. H. Raven and D. P. Gregory (1972[1973]) determined O. calcicola to be self-incompatible. It is known to form hybrids with O. suffrutescens. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Oenothera nutans is a PTH species and forms a ring of 14 chromosomes or a ring of 12 and 1 bivalent in meiosis, and is self-compatible and autogamous (W. Dietrich et al. 1997). It has plastome III and a BB genome composition. The disjunct occurrences of O. nutans in Arkansas and Missouri probably represent unintentional introductions by humans. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 10. | FNA vol. 10. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | Gaura calcicola | O. austromontana, O. biennis subsp. austromontana, O. biennis var. austromontana, O. biennis var. nutans |
Name authority | (P. H. Raven & D. P. Gregory) W. L. Wagner & Hoch: Syst. Bot. Monogr. 83: 211. (2007) | G. F. Atkinson & Bartlett: Rhodora 15: 83. (1913) |
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