Oenothera arizonica |
Oenothera xylocarpa |
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California evening primrose |
wood fruit evening primrose, woodyfruit evening primrose |
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Habit | Herbs winter-annual, younger parts sparsely to densely strigillose and sparsely to densely hirsute, older stems glabrate; from a taproot. | Herbs perennial, acaulescent, densely short-hirsute, also sometimes sparsely long-hirsute distally; from a thick, fleshy taproot. |
Stems | ascending to erect, with decumbent branches, thickened at base, tapering toward apex, 10–35(–60) cm. |
|
Leaves | in a basal rosette and cauline, basal 5–10(–26) × 0.6–1.5(–3.5) cm, cauline 5–8(–15.5) × 1–2 cm; petiole 0–12 cm; blade lanceolate to oblanceolate, margins pinnatifid or sometimes coarsely serrate. |
in a basal rosette, 2.6–4.2(–6.2) × 1.4–4.2 cm; petiole 2.5–9(–11.5) cm; blade usually oblanceolate to obovate, sometimes suborbiculate, margins dentate, pinnately lobed, lateral lobes oblong to lanceolate, often absent or reduced to only a few lobes toward terminal lobe, base rounded to cordate. |
Flowers | 1–several opening per day near sunset; buds nodding, weakly quadrangular, without free tips; floral tube 26–31 mm; sepals 19–26 mm, conspicuously maroon-spotted, each spot at base of a long hair; petals white, fading pink to deep pink, broadly obovate or obcordate, 16–26(–36) mm; filaments 9–15 mm, anthers 7–9 mm; style 45–50 mm, stigma exserted beyond anthers at anthesis. |
usually 1–3, rarely more, opening per day near sunset; buds erect, quadrangular, without free tips; floral tube 27–45(–55) mm; sepals 25–30 mm; petals intensely yellow, fading deep salmon red, obcordate, 25–38 mm; filaments 17–23 mm, anthers 7–10 mm; style 44–65(–80) mm, stigma somewhat exserted beyond anthers at anthesis. |
Capsules | spreading, woody in age, curved upward, or distal end recurved, cylindrical, obtusely 4-angled, especially toward base, tapering gradually from base to apex, 30–80 × 2.5–3.5 mm; sessile. |
moderately thin and flexible, lanceoloid, falcate or sigmoid, often contorted and twisted, 4-angled, 35–90 × 7–11 mm, gradually tapering to a long, slender, sterile apex, 10–30(–40) mm, valves conspicuously wrinkled, dehiscent 2/3–3/4 their length; sessile. |
Seeds | numerous, in 1 row per locule, light brown to yellowish brown with dark purple splotches, obovoid, 1.6–2 mm. |
numerous, in 1 row per locule, often forming 2 rows near base of capsule, obovoid, 2.4–3.2 × 1.3–1.7 mm, surface coarsely rugose. |
2n | = 14. |
= 14. |
Oenothera arizonica |
Oenothera xylocarpa |
|
Phenology | Flowering (Oct–)Feb–May. | Flowering Jun–Jul(–Aug). |
Habitat | Gravelly or sandy soil, along watercourses, disturbed sites. | Open meadows, flats or slopes on loose granitic gravel, sand, or pumice in Pinus jeffreyi forests with Artemisia tridentata, or in Pinus contorta subsp. murrayana and Abies magnifica forests. |
Elevation | 200–1400 m. (700–4600 ft.) | 2200–3100 m. (7200–10200 ft.) |
Distribution |
AZ; Mexico (Sonora) |
CA; NV |
Discussion | Oenothera arizonica occurs in southern Arizona from Maricopa and Yuma counties to Cochise County, and from scattered localities in northern Sonora, Mexico, including Cerro Tepopa, Puerto Libertad, and Tastiota. The populations from southwestern Arizona (Yuma County) southward to Sonora often grow on low dunes. Populations from sand dunes in Yuma County, Cabeza Prieta National Wildlife Refuge, appear to be a large phenotype of Oenothera arizonica that differ from all other specimens in the size of vegetative parts and flowers, and comprise all of the atypical measurements given in the description. Oenothera arizonica typically grows on dunes in Sonora, but rarely so in Arizona. Populations growing on dunes should be studied further and compared to non-dune populations in the northern and eastern portion of the range. Oenothera arizonica is self-compatible (W. L. Wagner et al. 2007; K. E. Theiss et al. 2010). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Oenothera xylocarpa is known from three disjunct areas in California and adjacent Nevada: Mount Rose, Washoe County, Nevada; southern Sierra Nevada, southwestern Mono County, California, from the vicinity of Crestview south to Casa Diablo; and the area in the southern Sierra Nevada bounded by Horseshoe and Big Whitney meadows to the east and north, and Casa Vieja and Volcano Meadows to the south and west, west-central Inyo and eastern Tulare counties, California. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 10. | FNA vol. 10. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | O. deltoides var. arizonica, O. avita subsp. arizonica, O. californica subsp. arizonica | Anogra xylocarpa |
Name authority | (Munz) W. L. Wagner: Novon 8: 308. (1998) | Coville: Contr. U.S. Natl. Herb. 4: 105, plate 8. (1893) |
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