Oenothera argillicola |
Oenothera mckelveyae |
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shale barren evening primrose |
Mckelvey's beeblossom |
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Habit | Herbs biennial or short-lived perennial, strigillose and sparsely to moderately villous, hairs sometimes pustulate, pustules with green or red bases, inflorescence glabrous or sparsely glandular puberulent, sometimes also sparsely villous. | Herbs perennial, clumped, long-villous, more sparsely so distally, hairs erect, 2–4 mm, also strigillose, sometimes glabrate distally or also sparsely glandular puberulent; from twisted, woody rootstock. |
Stems | erect to ascending, green or red, unbranched or with branches obliquely arising from rosette or in distal 1/2 of main stem. |
ascending, branched below or just above ground, branched also proximal to inflorescences, 30–70(–120) cm. |
Leaves | in a basal rosette and cauline, basal 7–25 × 0.7–2 cm, cauline 6–13 × 0.4–1 cm; blade dark green, somewhat glossy, very narrowly oblanceolate to narrowly oblanceolate, linear-elliptic, lanceolate, or nearly linear, margins flat, entire or remotely and bluntly dentate, sometimes with larger teeth near base; bracts persistent. |
in a basal rosette and cauline, basal 3–17 × 0.8–2 cm, blade oblanceolate, cauline 1–6.5 × 0.1–1.5 cm, sessile, blade narrowly oblanceolate to elliptic, margins conspicuously sinuate-dentate, often undulate. |
Inflorescences | curved with ascending tip, unbranched. |
slender. |
Flowers | opening near sunset; buds erect, 4–8 mm diam., with free tips subterminal, divergent and hornlike, 3–9 mm; floral tube 32–52 mm; sepals yellowish green to yellow, sometimes flushed with red, especially at apex, 27–38 mm; petals yellow to pale yellow, fading pale yellow to pale yellowish orange, very broadly obcordate or obovate, 25–42 mm; filaments 20–27 mm, anthers 9–13 mm, pollen 90–100% fertile; style 60–85 mm, stigma exserted beyond anthers at anthesis. |
4-merous, zygomorphic, opening near sunset; floral tube 2–3.5 mm; sepals 6–12 mm; petals white, fading dark pink to red, slightly unequal, elliptic-obovate, 7–11 mm, long-clawed; stamens presented in lower 1/2 of flower, filaments 5–9 mm, lanate at very base, anthers 2–4 mm, pollen 90–100% fertile; style 9–16 mm, stigma exserted beyond anthers at anthesis. |
Capsules | spreading at nearly a right angle to stem, curved upward, sometimes secund, dull green or rusty brown when dry, narrowly lanceoloid to lanceoloid, 20–40 × 4–6 mm, free tips of valves 1–2 mm. |
reflexed, lanceoloid to narrowly ovoid, narrowly 4-winged, 8–19 × 1.5–2 mm, tapering to a sterile stipe 3–9 mm. |
Seeds | 1.3–1.9 × 0.7–1.1 mm. |
(1 or)2–4, 2–3 × 1 mm, yellowish to reddish brown. |
2n | = 14. |
= 14. |
Oenothera argillicola |
Oenothera mckelveyae |
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Phenology | Flowering (Jun–)Jul–Oct. | Flowering Mar–Jun. |
Habitat | Open sites on Devonian Brallier shale slopes, barrens, outcrops or adjacent roadsides in mid-Appalachian Allegheny Mountains. | Sandy soil. |
Elevation | 150–700 m. (500–2300 ft.) | 0–300 m. (0–1000 ft.) |
Distribution |
MD; PA; VA; WV |
TX; Mexico (Nuevo León, Tamaulipas) |
Discussion | Oenothera argillicola is one of eight angiosperm species restricted to the Devonian Brallier shale barrens, but among them only O. argillicola and Trifolium virginicum occur throughout the shale barren region. Oenothera argillicola has plastome V and a CC genome composition. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Oenothera mckelveyae, on the Rio Grande Plain, is found in an area bounded by from Dimmit and LaSalle counties east to Karnes and Refugio counties in the north, southward through south Texas, extending to northeastern Tamaulipas and adjacent Nuevo León. P. H. Raven and D. P. Gregory (1972[1973]) found Oenothera mckelveyae to be self-incompatible. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 10. | FNA vol. 10. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | O. argillicola var. pubescens | Gaura villosa var. mckelveyae, Gaura mckelveyae |
Name authority | Mackenzie: Torreya 4: 56. (1904) | (Munz) W. L. Wagner & Hoch: Syst. Bot. Monogr. 83: 213. (2007) |
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