Oenothera acutissima |
Oenothera nutans |
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flaming Gorge evening primrose |
nodding evening-primrose |
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Habit | Herbs perennial, subacaulescent or very short-caulescent, strigillose mostly along leaf margins and flower parts, also sparsely glandular puberulent, sometimes also sparsely hirsute distally; from a stout taproot, usually with several long, lateral roots often producing adventitious shoots. | Herbs biennial, often appearing glabrous to naked eye, usually strigillose and sparsely to moderately villous proximal to inflorescence, hairs translucent and with or without pustules, pustules not red (in fresh material), inflorescence glabrous, glandular puberulent, or strigillose and glandular puberulent. |
Stems | (when present) ascending, (1–)several–10, densely leafy, 1–2 cm. |
erect, red on proximal parts, green on distal ones, rarely red throughout, unbranched or with branches obliquely arising from rosette and secondary branches arising from main stem, 30–200 cm. |
Leaves | primarily in a basal rosette, 7–14(–18) × (0.3–)0.5–1(–1.5) cm, moderately thick and stiff; petiole (1.2–)3–5 cm; blade linear to very narrowly elliptic, margins irregularly and coarsely dentate or pinnately lobed, apex long-attenuate. |
in a basal rosette and cauline, basal 10–32 × 3–7 cm, cauline 6–20 × 2–8 cm; blade green to pale green, narrowly oblanceolate to narrowly obovate, or narrowly elliptic, sometimes lanceolate distally, margins usually flat, rarely undulate, bluntly dentate, teeth widely spaced, sometimes sinuate-dentate proximally; bracts caducous. |
Inflorescences | erect, unbranched or with secondary branches just proximal to main one. |
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Flowers | 1–3 opening per day near sunset; buds with unequal free tips 1–3 mm; floral tube (53–)60–100 mm; sepals 26–50 mm; petals bright yellow, fading deep reddish orange, drying purplish brown, 28–50 mm; filaments 21–35 mm, anthers 9–11 mm; style 75–143 mm, stigma exserted beyond anthers. |
opening near sunset; buds erect, 4–6 mm diam., with free tips terminal, erect, 1.5–6 mm; floral tube 30–43 mm; sepals yellowish green or flushed with red, 10–23 mm; petals yellow to pale yellow, fading pale yellowish white, very broadly obcordate, 14–25(–30) mm; filaments 10–25 mm, anthers 4–10 mm, pollen ca. 50% fertile; style 35–63 mm, stigma surrounded by anthers at anthesis. |
Capsules | leathery in age, oblong-oblanceoloid, narrowly winged, wings oblong, 1–2(–4) mm wide, broadest near apex, 14–18(–22) × 7–8 mm (excluding wings), apex abruptly constricted, dehiscent 1/4–1/3 their length, valve surface with inconspicuous veins; sessile. |
erect or slightly spreading, dull green when dry, narrowly lanceoloid to narrowly ovoid, 12–36 × 3–6 mm, free tips of valves 1–1.5 mm. |
Seeds | asymmetrically cuneiform, 2–2.5 mm. |
1–1.9 × 0.6–0.9 mm. |
2n | = 14. |
= 14. |
Oenothera acutissima |
Oenothera nutans |
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Phenology | Flowering May–Jun. | Flowering Jun–Aug(–Sep). |
Habitat | Restricted to sandy and gravelly, reddish, soil in seasonally wet sites, meadows, depressions, along arroyos, among rocks, in mixed conifer forests, sagebrush scrub. | Open, often disturbed sites, stream beds, flood plains, slopes, margins of mixed deciduous forests, roadsides, old fields. |
Elevation | 1800–2400(–2600) m. (5900–7900(–8500) ft.) | (200–)400–1700 m. ((700–)1300–5600 ft.) |
Distribution |
CO; UT |
AL; AR; CT; DE; FL; GA; IN; KY; MA; MD; ME; MI; MO; NC; NH; NJ; NY; OH; PA; SC; TN; VA; VT; WV; ON |
Discussion | Oenothera acutissima is known only from the vicinity of Manila, eastern Uinta Mountains, Daggett and Duchesne counties, Utah, east to areas in and near the foothills of the Douglas and Blue mountains, in Uinta County, Utah, and Moffat County, Colorado. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Oenothera nutans is a PTH species and forms a ring of 14 chromosomes or a ring of 12 and 1 bivalent in meiosis, and is self-compatible and autogamous (W. Dietrich et al. 1997). It has plastome III and a BB genome composition. The disjunct occurrences of O. nutans in Arkansas and Missouri probably represent unintentional introductions by humans. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 10. | FNA vol. 10. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | O. flava var. acutissima | O. austromontana, O. biennis subsp. austromontana, O. biennis var. austromontana, O. biennis var. nutans |
Name authority | W. L. Wagner: Syst. Bot. 6: 153, fig. 1. (1981) | G. F. Atkinson & Bartlett: Rhodora 15: 83. (1913) |
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