Noccaea fendleri subsp. siskiyouensis |
Noccaea fendleri |
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Siskiyou penny-cress |
alpine pennycress, Fendler's pennycress, penny-cress, wild candytuft |
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Habit | Perennials; (cespitose or not). | |||||||||||||||||
Stems | (0.6–)0.7–1.6(–2.4) dm. |
simple or several from caudex, erect or ascending, unbranched or branched distally, (0.1–)0.4–3.2(–4.5) dm. |
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Basal leaves | petiole differentiated from blade, (1.7–)2–3.6(–4) times longer than blade; blade ovate to oblong, 4–8(–9) mm wide, base cuneate. |
petiole 0.4–7.3 cm; blade linear, oblong, oblanceolate, ovate, obovate, or spatulate, 0.4–3 cm × 2–20 mm, base cuneate to attenuate, margins entire, denticulate, or dentate, apex obtuse to acute. |
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Cauline leaves | 2–6. |
2–21; blade ovate or suboblong, 0.4–2.8 × 0.1–1.7 mm, base auriculate to subamplexicaul, margins entire or dentate, apex obtuse to subacute. |
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Racemes | lax to compact. |
0.5–25 cm, (congested or considerably elongated). |
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Flowers | petals white, 3.8–6 × 1.2–2.2(–2.9) mm. |
sepals 1.6–5.3 × 0.5–1.5 mm, (margins membranous); petals white to pinkish purple, (3.4–)4.2–13 × 1–4.9 mm, often flaring between blade and claw, apex obtuse; filaments 2–7.5 mm; anthers 0.5–1 mm. |
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Fruiting pedicels | horizontal or slightly descending, forming an angle 80º or more with rachis. |
horizontal to ascending, straight, 2.5–15 mm. |
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Fruits | often strongly winged, obovate to obdeltate, (4–)4.8–7(–8) × (3–)3.5–5.5(–6) mm, 1–2 times as long as wide, apex truncate to strongly emarginate; style (0.8–)1–2 mm. |
obovate, obcordate, obdeltate, or elliptic, 2.5–16 × 1.5–9 mm, usually winged, rarely not winged apically, base cuneate, apex obtuse, truncate, or emarginate; ovules 4–10 per ovary; style (0.4–)1.1–4.2 mm. |
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Seeds | 2–4 per fruit. |
brown, 1.1–2.1 mm, minutely reticulate. |
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2n | = 14. |
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Noccaea fendleri subsp. siskiyouensis |
Noccaea fendleri |
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Phenology | Flowering Apr–Jun. | |||||||||||||||||
Habitat | Moist, open rocky serpentine slopes | |||||||||||||||||
Elevation | 300-500 m [1000-1600 ft] | |||||||||||||||||
Distribution |
OR |
AZ; CA; CO; ID; MT; NM; NV; OR; TX; UT; WA; WY; n Mexico
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Discussion | Subspecies siskiyouensis is known from Curry, Douglas, and Josephine counties. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Subspecies 5 (5 in the flora). Some North American authors (e.g., P. K. Holmgren 1971; R. C. Rollins 1993) treated Noccaea fendleri as conspecific with the European Thlaspi montanum, whereas others (e.g., E. B. Payson 1926) recognized the North American taxa as endemic to the continent. Molecular studies (M. Koch and I. A. Al-Shehbaz 2004) supported Payson’s view of the independent status of the North American representatives. Although Payson and Rollins recognized more than one species here (e.g., T. californicum, T. idahoense, T. glaucum, T. fendleri), I agree with Holmgren in treating them as infraspecific taxa, but at the subspecific instead of varietal rank. The distinctions among them are not very sharp, and hybridization appears to have played some role in blurring their boundaries, as evidenced from Holmgren’s report of morphologically intermediate forms between subsp. glauca (as T. montanum var. montanum) and each of subspp. fendleri, idahoensis, and siskiyouensis. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 7, p. 603. | FNA vol. 7, p. 601. | ||||||||||||||||
Parent taxa | ||||||||||||||||||
Sibling taxa | ||||||||||||||||||
Subordinate taxa | ||||||||||||||||||
Synonyms | Thlaspi montanum var. siskiyoense | Thlaspi fendleri, Thlaspi montanum var. fendleri | ||||||||||||||||
Name authority | (P. K. Holmgren) Al-Shehbaz & M. Koch: Syst. Bot. 29: 383. (2004) | (A. Gray) Holub: Preslia 70: 108. (1998) | ||||||||||||||||
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