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Avery's sword fern, swordfern

Boston fern, narrow swordfern, sword fern, tuber ladder fern, tuber sword fern, tuberous sword fern

Stem

scales loosely appressed to spreading, essentially concolored.

scales spreading, concolored.

Leaves

8–30 × 0.5–1.3 dm.

2.5–10.7 × 0.3–0.7 dm.

Petiole

1.2–5 dm, moderately to densely scaly;

scales spreading, reddish to light brown throughout.

0.3–2 dm, moderately to densely scaly;

scales spreading, pale brown throughout.

Blade

moderately scaly, glabrous to occasionally pubescent abaxially, glabrous adaxially, hairs pale to light brown, 0.2–0.7 mm.

lacking scales, glabrous (rarely with a few branched hairs abaxially).

Indusia

horseshoe-shaped to circular, attached at narrow sinus or peltate, 0.9–1.1 mm wide.

reniform to lunate or deltate-rounded, attached along broad sinus, 1.1–1.7 mm wide.

Tubers

absent.

present or absent.

Rachis

5–26 dm, points of pinna attachment 9–25 mm apart;

scales scattered, brown throughout.

2.2–9 dm, points of pinna attachment 5–12 mm apart;

scales moderately spaced to dense, pale to dark brown, point of attachment distinctly darker.

Central

pinnae narrowly deltate to oblong-deltate, 3.6–9 × 0.7–2 cm, base truncate, sometimes rounded basiscopically, barely auriculate acroscopically, acroscopic lobe obtuse to acute, margins serrulate, apex narrowly acute to attenuate, falcate to slightly so;

costae adaxially sparsely pubescent, hairs pale, erect, 0.1–0.5 mm.

pinnae oblong to lanceolate-oblong, straight to slightly falcate, 0.9–5 × 0.4–0.9 cm, base auriculate-cordate, acroscopically overlapping rachis, acroscopic lobe deltate, margins entire to serrulate to smoothly crenate, apex acute to bluntly rounded;

costae adaxially glabrous.

2n

= 164.

= 82.

Nephrolepis ×averyi

Nephrolepis cordifolia

Habitat Terrestrial or epiphytic in forested, often moist habitats, e.g., swamps, hammocks, or relatively open, disturbed habitats Terrestrial or epiphytic in wet, shady places, limestone ledges, cliffs, rock, roadsides, and often old homesites or waste places, widely escaped from cultivation and only questionably native to any particular region
Elevation 0 m (0 ft) 0 m (0 ft)
Distribution
from FNA
FL; Mexico; West Indies
[BONAP county map]
from FNA
FL; HI; Mexico; Central America; South America; West Indies; Africa; se Asia
[WildflowerSearch map]
[BONAP county map]
Discussion

Nephrolepis × averyi is an allotetraploid derived from hybridization between N. biserrata and N. exaltata and is known to occur only in mixed populations of both parent species. It is distinguished from N. exaltata by its sparsely hairy adaxial costae, larger size, and misshapen spores, and from N. biserrata by its falcate pinnae and narrower leaves.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 2. FNA vol. 2.
Parent taxa Dryopteridaceae > Nephrolepis Dryopteridaceae > Nephrolepis
Sibling taxa
N. biserrata, N. cordifolia, N. exaltata, N. multiflora
N. biserrata, N. exaltata, N. multiflora, N. ×averyi
Synonyms Polypodium cordifolium, Aspidium cordifolium
Name authority Nauman: Amer. Fern J. 69: 69. (1979) (Linnaeus) C. Presl: Tent. Pterid. 79. (1836)
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