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serrated tussock, serrated tussockgrass, yass tussockgrass

nassella, needlegrass, tussockgrass

Habit Plants perennial; cespitose, not rhizomatous. Plants usually perennial, rarely annual; cespitose, occasionally rhizomatous.
Culms

20-60 cm tall, about 1 mm thick, erect, internodes glabrous;

nodes 2-4, pubescent.

10-175(210) cm, sometimes branched at the upper nodes, branches flexible;

prophylls not evident, shorter than the sheaths.

Sheaths

glabrous, smooth;

collars glabrous, without tufts of hair at the sides;

ligules 0.5-2.5 mm, glabrous, obtuse;

blades 15-45 cm long, 0.2-0.6 mm wide, convolute, stiff, scabridulous.

Leaves

mostly basal, not overwintering;

sheaths open;

cleistogenes sometimes present;

auricles absent;

ligules membranous, sometimes pubescent or ciliate;

blades of basal leaves 3-60 cm long, 0.2-8 mm wide, apices narrowly acute to acute, not sharp, flag leaf blades 1-80 mm, bases about as wide as the top of the sheaths.

Panicles

8-25 cm, open, lax, sparse;

branches 2-6 cm, with 1-8 spikelets;

pedicels 3-12 mm.

Inflorescences

terminal panicles, sometimes partially included at maturity.

Spikelets

3-22 mm, with 1 floret;

rachillas not prolonged beyond the base of the floret;

disarticulation above the glumes, beneath the floret.

Glumes

subequal, 4-10 mm long, 0.9-1.2 mm wide, narrowly lanceolate, scabridulous, apices aristate;

florets 1.5-2.5 mm long, 0.7-0.9 mm wide, terete, widest near the top;

calluses 0.1-0.3 mm, acute, strigose, hairs reaching to midlength on the lemmas;

lemmas strongly tuberculate, particularly distally, mostly glabrous, narrowing abruptly to the crown, midveins pubescent proximally;

crowns about 0.5 mm long, about 0.2 mm wide, straight-sided, rims entire or irregularly lacerate, glabrous;

awns 15-35 mm, eccentric, straight to twice-geniculate;

anthers 3, 1-1.5 mm.

longer than the floret, narrowly lanceolate or ovate, basal portion usually purplish at anthesis, color fading with age, (1)3-5-veined, sometimes awned;

florets usually terete, sometimes slightly laterally compressed;

calluses blunt or sharp, glabrous or antrorsely strigose;

lemmas usually papillose or tuberculate, at least distally, sometimes smooth throughout, glabrous or variously hairy, strongly convolute, wrapping 1.2-1.5 times around the caryopses, apices not lobed, fused distally into crowns, these often evident by their pale color and constricted bases;

crowns mostly glabrous, rims often bearing hairs with bulbous bases;

awns terminal, centric or eccentric, deciduous or persistent, usually twice-geniculate, second geniculation often obscure;

paleas up to 1/2 as long as the lemmas, glabrous, without veins, flat;

lodicules 2 or 3, if 3, the third somewhat shorter than the other 2;

anthers 1 or 3, if 3, often of 2 lengths, penicillate;

ovaries glabrous;

styles 2, bases free.

Caryopses

about 1.2 mm, oblong, dark brown.

glabrous, not ribbed;

hila elongate;

embryos to 2/5 as long as the caryopses.

x

= 7, 8.

2n

= 36.

Nassella trichotoma

Nassella

Distribution
from FNA
IL; KY; NC; SC
[BONAP county map]
from FNA
AL; AR; AZ; CA; CO; IA; ID; IL; KS; KY; LA; MN; MT; NC; ND; NE; NM; NY; OK; OR; SC; SD; TX; UT; WI; WY; HI; AB; BC; MB; NT; SK; YT
[BONAP county map]
Discussion

Nassella trichotoma is a native of South America, and has been accidentally introduced into the United States. Because it is on the U.S. Department of Agriculture's noxious weed list, all known populations have been eliminated. New populations should be reported to the Department.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Nassella used to be interpreted as a South American genus of approximately 14 species. It is now interpreted as including at least 116 species (Barkworth and Torres 2001), the majority of which are South American. The additional species were previously included in Stipa. There are eight species in the Flora region, one of which is introduced; two additional species treated here were found in the region at one time, but have not become established. The strongly convolute lemmas distinguish Nassella from all other genera of Stipeae in the Americas and, in combination with the reduced, ecostate, glabrous paleas, from all other genera in the tribe worldwide. Molecular data (Jacobs et al. 2006) support the expanded interpretation of Nassella. Relationships among the species have not been explored.

Many species of Nassella develop both cleistogamous and chasmogamous florets in the terminal panicle. The cleistogamous florets have 1-3 anthers that are less than 1 mm long; the chasmogamous florets have 3 anthers that are significantly longer. In addition, some species develop panicles in the axils of their basal sheaths. Spikelets of cleistogenes have reduced or no glumes, and florets with no or very short awns.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Florets 1.5-3 mm long; blades 0.2-1.5 mm wide, usually tightly convolute.
→ 2
2. Florets widest about midlength; awns 45-100 mm long, almost centric
N. tenuissima
2. Florets widest near the top; awns 7-35 mm long, eccentric.
→ 3
3. Awns 15-35 mm long; lemmas strongly tuberculate, particularly distally
N. trichotoma
3. Awns 7-10 mm long; lemmas smooth
N. chilensis
1. Florets 3.4-13 mm long; blades 0.4-8 mm wide, flat to convolute.
→ 4
4. Terminal segment of the awns cernuous.
→ 5
5. Awns 12-55 mm long, 0.1-0.2 mm thick at the base
N. lepida
5. Awns 50-110 mm long, 0.2-0.3 mm thick at the base
N. cemua
4. Terminal segment of the awns straight.
→ 6
6. Florets 3.4-5.5 mm long; lemmas not constricted below the crown; awns 19-32 mm long
N. viridula
6. Florets 6-13 mm long; lemmas constricted below the crown; awns 30-120 mm long.
→ 7
7. Lemmas hairy between the veins at maturity
N. pulchra
7. Lemmas glabrous between the veins at maturity.
→ 8
8. Crowns usually wider than long, the rims with hairs to 0.5 mm long; florets widest just below the crowns
N. neesiana
8. Crowns usually longer than wide, the rims with hairs 1-2 mm long; florets widest near or slightly above midlength.
→ 9
9. Florets 6.5-13 mm long; crowns often flaring distally; plants native to Texas and adjacent states
N. leucotricha
9. Florets 6-8 mm long; crowns more or less straight-sided; plants introduced, established in California
N. manicata
Source FNA vol. 24, p. 177. FNA vol. 24, p. 170. Author: Mary E. Barkworth;.
Parent taxa Poaceae > subfam. Pooideae > tribe Stipeae > Nassella Poaceae > subfam. Pooideae > tribe Stipeae
Sibling taxa
N. cernua, N. chilensis, N. lepida, N. leucotricha, N. manicata, N. neesiana, N. pulchra, N. tenuissima, N. viridula
Subordinate taxa
N. cemua, N. chilensis, N. lepida, N. leucotricha, N. manicata, N. neesiana, N. pulchra, N. tenuissima, N. trichotoma, N. viridula
Name authority (Nees) Hack, ex Arechav. (Trin.) E. Desv.
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