Nassella neesiana |
Nassella |
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Uruguayan needlegrass, Uruguayan tussockgrass |
nassella, needlegrass, tussockgrass |
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Habit | Plants perennial; cespitose, not rhizomatous. | Plants usually perennial, rarely annual; cespitose, occasionally rhizomatous. | ||||||||||||||||||||||||||||||||||||
Culms | 30-140 cm tall, 1-1.8 mm thick, erect or geniculate, internodes glabrous; nodes usually 2-4, exposed, sericeous, hairs to 1.2 mm. |
10-175(210) cm, sometimes branched at the upper nodes, branches flexible; prophylls not evident, shorter than the sheaths. |
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Sheaths | glabrous or slightly hispid, basal leaf sheaths often with cleistogenes; collars glabrous, often brown or purple-tinged, with tufts of hair at the sides, hairs 0.5-1.2 mm; Iigules 1-4 mm, glabrous or pubescent, truncate; blades to 30 cm long, 2-8 mm wide, flat to convolute, sometimes scabrous, sometimes with hairs. |
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Leaves | mostly basal, not overwintering; sheaths open; cleistogenes sometimes present; auricles absent; ligules membranous, sometimes pubescent or ciliate; blades of basal leaves 3-60 cm long, 0.2-8 mm wide, apices narrowly acute to acute, not sharp, flag leaf blades 1-80 mm, bases about as wide as the top of the sheaths. |
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Inflorescences | terminal panicles, sometimes partially included at maturity. |
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Spikelets | 3-22 mm, with 1 floret; rachillas not prolonged beyond the base of the floret; disarticulation above the glumes, beneath the floret. |
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Glumes | subequal, 10-22 mm long, 1.8-2.3 mm wide, narrowly lanceolate, glabrous, 3-5-veined; florets 6-13 mm long, 1-1.5 mm wide, terete, widest just below the crown; calluses 2-4.5 mm, sharp, strigose; lemmas often purple, finely rugose-papillose, particularly near the crown, constricted below the crown, midveins pilose proximally, glabrous between the veins at maturity; crowns 0.4-1.6 mm, usually wider than long, sides usually flaring somewhat distally, rims with hairs to 0.5 mm; awns 50-120 mm, clearly twice-geniculate, terminal segment straight; anthers 3-3.5 mm, penicillate. |
longer than the floret, narrowly lanceolate or ovate, basal portion usually purplish at anthesis, color fading with age, (1)3-5-veined, sometimes awned; florets usually terete, sometimes slightly laterally compressed; calluses blunt or sharp, glabrous or antrorsely strigose; lemmas usually papillose or tuberculate, at least distally, sometimes smooth throughout, glabrous or variously hairy, strongly convolute, wrapping 1.2-1.5 times around the caryopses, apices not lobed, fused distally into crowns, these often evident by their pale color and constricted bases; crowns mostly glabrous, rims often bearing hairs with bulbous bases; awns terminal, centric or eccentric, deciduous or persistent, usually twice-geniculate, second geniculation often obscure; paleas up to 1/2 as long as the lemmas, glabrous, without veins, flat; lodicules 2 or 3, if 3, the third somewhat shorter than the other 2; anthers 1 or 3, if 3, often of 2 lengths, penicillate; ovaries glabrous; styles 2, bases free. |
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Caryopses | 3-5 mm. |
glabrous, not ribbed; hila elongate; embryos to 2/5 as long as the caryopses. |
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Terminal | panicles 5-40 cm, exserted, erect to nodding; branches 2.5-8.5 cm, with 2-5 spikelets; pedicels 1-8 mm, angled, scabrous, pubescent. |
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x | = 7, 8. |
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2n | = 28. |
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Nassella neesiana |
Nassella |
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Distribution |
AL |
AL; AR; AZ; CA; CO; IA; ID; IL; KS; KY; LA; MN; MT; NC; ND; NE; NM; NY; OK; OR; SC; SD; TX; UT; WI; WY; HI; AB; BC; MB; NT; SK; YT |
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Discussion | Nassella neesiana is native to South America, growing from Ecuador to Argentina, primarily in steppe habitats. It was found on ballast dumps in Mobile, Alabama but has not persisted in the Flora region. It has become established in Australia, where it is considered a noxious weed. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Nassella used to be interpreted as a South American genus of approximately 14 species. It is now interpreted as including at least 116 species (Barkworth and Torres 2001), the majority of which are South American. The additional species were previously included in Stipa. There are eight species in the Flora region, one of which is introduced; two additional species treated here were found in the region at one time, but have not become established. The strongly convolute lemmas distinguish Nassella from all other genera of Stipeae in the Americas and, in combination with the reduced, ecostate, glabrous paleas, from all other genera in the tribe worldwide. Molecular data (Jacobs et al. 2006) support the expanded interpretation of Nassella. Relationships among the species have not been explored. Many species of Nassella develop both cleistogamous and chasmogamous florets in the terminal panicle. The cleistogamous florets have 1-3 anthers that are less than 1 mm long; the chasmogamous florets have 3 anthers that are significantly longer. In addition, some species develop panicles in the axils of their basal sheaths. Spikelets of cleistogenes have reduced or no glumes, and florets with no or very short awns. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 24, p. 172. | FNA vol. 24, p. 170. | ||||||||||||||||||||||||||||||||||||
Parent taxa | Poaceae > subfam. Pooideae > tribe Stipeae > Nassella | Poaceae > subfam. Pooideae > tribe Stipeae | ||||||||||||||||||||||||||||||||||||
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Synonyms | Stipa neesiana | |||||||||||||||||||||||||||||||||||||
Name authority | (Trin. & Rupr.) Barkworth | (Trin.) E. Desv. | ||||||||||||||||||||||||||||||||||||
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