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bayberry, myrique, sweet gale, wax-myrtle

southern bayberry, southern wax-myrtle

Habit Shrubs or small trees, often aromatic and resinous. Shrubs or small trees, evergreen, often forming large, rhizomatous colonies of much-branched specimens, to 14 m.
Branches

spreading, terete, glabrous or pubescent, often gland-dotted.

Branchlets

reddish brown, densely gland-dotted when young, otherwise glabrous to densely pilose, eventually glabrate;

glands yellow.

Leaves

blade aromatic when crushed (except M. inodora), oblanceolate, elliptic, obovate, or oblong-ovate, membranous or leathery, margins entire or serrate-denticulate, especially in distal 1/2, pubescent or glabrous, usually gland-dotted.

blade aromatic when crushed, linear-oblanceolate to obovate, (1.1-)2-10.5(-13.3) × 0.4-3.3 cm, leathery, base cuneate to attenuate, margins entire or coarsely serrate beyond middle, apex acute to slightly rounded;

surfaces abaxially pale yellow-green, glabrous except for pilose midrib, adaxially dark green, glabrous to pilose, both surfaces densely glandular;

glands yellow to orange.

Inflorescences

± erect, ellipsoid to short-cylindric or ovoid, appearing before or with leaves;

bracts ovate, glabrous or variously pubescent.

staminate 0.4-1.9 cm;

pistillate 0.3-1.5 cm.

Flowers

unisexual, rarely bisexual, staminate and pistillate flowers usually on different plants, infrequently on same plants.

unisexual, staminate and pistillate on different plants.

Staminate flowers

stamens (2-)3-12(-22), shorter or longer than subtending bract;

filaments mostly distinct, often connate into branching staminal column, each branch terminated by anther;

rudimentary ovary occasionally present.

bract of flower shorter than staminal column, margins opaque, densely ciliate, abaxially densely gland-dotted;

stamens mostly 3-4.

Pistillate flowers

ovary subtended by 2-6 broadly ovate bracteoles, these sometimes persistent and accrescent, always shorter than fruit, sometimes completely absent;

styles short.

bracteoles persistent in fruit, 4, not accrescent or adnate to fruit wall, margins ciliate, abaxially densely gland-dotted;

ovary glandular, especially at apex near style base.

Fruits

globose or ovoid to lenticular, smooth or more commonly with warty protuberances, usually covered with waxy coating that dries white.

globose-ellipsoid, 2-3.5(-4) mm;

fruit wall glabrous or sparsely glandular when young, obscured by enlarged protuberances and thick coat of blue-white wax.

x

= 8.

Myrica

Myrica cerifera

Phenology Flowering mid winter–spring, fruiting summer–fall.
Habitat Bogs, edges of marshes, ponds, creeks, and swamps, pine forests, mixed deciduous forests, pine barrens, coastal sand dunes, open fields, sandy hillsides
Elevation 0-450 m [0-1500 ft]
Distribution
map from USDA
Nearly worldwide
[BONAP county map]
map from FNA
AL; AR; DE; FL; GA; LA; MD; MS; NC; NJ; OK; SC; TX; VA; Mexico; Central America; West Indies; Bermuda
Discussion

Species ca. 50 (7 in the flora).

Myrica is often cultivated. Myrica species were used by various tribes of Native Americans for medicinal purposes. Leaves were used for a gynecological aid and an emetic; the bark, as a blood purifier and a kidney aid (D. E. Moerman 1986). Bayberry candles were used by early settlers, and they remain popular household items, both decorative and functional.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Myrica cerifera is an extremely variable species with respect to habitat and corresponding habit/vegetative morphology. In general, plants that occupy dry, sandy (more xeric) areas tend to be strongly rhizomatous, colonial, and smaller in stature, and to possess smaller leaves (commonly recognized as M. cerifera var. pumila). In contrast, plants of more mesic areas are seldom rhizomatous, not colonial, and often large and treelike, and they have larger leaves. These "extremes pass insensibly into each other" (J. W. Thieret 1966). I agree with Thieret's contention that these differences do not constitute reliable criteria upon which one should base taxonomic distinctions. Until it can be determined with certainty whether these differences are due to genetics or environment, the question will remain open. I have chosen the conservative route.

Myrica cerifera has often been confused with M. pensylvanica and with M. heterophylla. It is distinguished from M. pensylvanica on the basis of gland density on the leaves, the presence of glands versus hirsute pubescence on the fruit wall and protuberances (especially visible on young fruits), and less reliably on the size of the fruit (2-3.5 versus 3.5-5.5 mm). Myrica cerifera is distinguished from M. heterophylla by the density of the glands on the leaves and the glandular versus glabrous (usually) fruit wall.

Native Americans used a decoction of the leaves and stems of Myrica cerifera to treat fevers; and roots, to treat inflamed tonsils and stomachaches, and as a stimulant (D. E. Moerman 1986).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Parent taxa Myricaceae Myricaceae > Myrica
Sibling taxa
M. californica, M. gale, M. hartwegii, M. heterophylla, M. inodora, M. pensylvanica
Subordinate taxa
M. californica, M. cerifera, M. gale, M. hartwegii, M. heterophylla, M. inodora, M. pensylvanica
Key
1. Bracts of staminate flowers longer than stamens; bracteoles of pistillate flowers 2, accrescent and adnate to fruit; fruits smooth (without protuberances), lacking waxy deposit (subg. Myrica).
→ 2
1. Bracts of staminate flowers shorter than stamens; bracteoles of pistillate flowers 4–6, not accrescent or adnate to fruit; fruits with numerous protuberances, usually covered with waxy coating that dries white or blue-white to gray (subg. Morella Loureiro).
→ 3
2. Bracteoles of pistillate flowers glabrous; leaf margins serrate, usually minutely so, with 1–4 pairs of teeth restricted to distal 1/3 of blade; not extending s of Oregon in w part of range.
M. gale
2. Bracteoles of pistillate flowers densely pilose, especially at apex; leaf margins serrate, often coarsely so, with 4–12 pairs of teeth ± in distal 1/2 of blade; California only.
M. hartwegii
3. Staminate flowers with 6 or more stamens, rarely 2–3, especially in distal flowers; fruit wall, but not warty protuberances, pubescent.
→ 4
3. Staminate flowers with 3–5(–7) stamens; fruit wall usually glabrous, if pubescent, warty protuberances also pubescent.
→ 5
4. Margins of leaf blade entire, rarely serrate at apex; blade not aromatic when crushed; flowers unisexual, staminate and pistillate on different plants; restricted to Gulf Coast region.
M. inodora
4. Margins of leaf blade conspicuously serrate almost their entire length; blade fragrant when crushed; flowers bisexual, staminate, and pistillate, all on same plant; Pacific Coast region.
M. californica
5. Leaf blade densely glandular on both surfaces.
M. cerifera
5. Leaf blade densely glandular only abaxially.
→ 6
6. Fruit wall and warty protuberances densely hirsute when young; branches whitish gray in age; leaves deciduous, membranous; fruits 3.5–5.5 mm.
M. pensylvanica
6. Fruit wall glabrous or sparsely glandular, warty protuberances ± glandular; branches black; leaves persistent or tardily deciduous, leathery; fruits 3–4.5 mm.
M. heterophylla
Synonyms Cerothamnus, Gale, Morella Cerophora lanceolata, Cerothamnus arborescens, Cerothamnus ceriferus, Cerothamnus pumilus, Morella cerifera, M. cerifera var. angustifolia, M. cerifera var. arborescens, M. cerifera var. dubia, M. cerifera var. pumila, M. pumila, M. pusilla
Name authority Linnaeus: Sp. Pl. 2: 1024. 175: Gen. Pl. ed. 5, 449. (1754) Linnaeus: Sp. Pl. 2: 1024. (1753)
Source FNA vol. 3. Treatment author: Allan J. Bornstein. FNA vol. 3. Treatment author: Allan J. Bornstein.
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