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nimblewell, nimblewill, nimblewill muhly

canyon muhly, muhly

Habit Plants perennial (appearing annual); usually cespitose, not rhizomatous, sometimes stoloniferous. Plants perennial; cespitose, not rhizomatous.
Culms

10-45(70) cm, geniculate, often rooting at the lower nodes, glabrous or puberulent below the nodes;

internodes often smooth, shiny, glabrous.

60-140 cm, erect;

internodes puberulent or glabrous for most of their length, puberulent below the nodes.

Sheaths

shorter than the internodes, glabrous for most of their length, margins shortly (0.3-1.2 mm) pubescent distally, not becoming spirally coiled when old;

ligules 0.2-0.5 mm, truncate, erose, ciliate;

blades (1)3-10 cm long, 1-4.5 mm wide, flat, smooth or scabridulous.

shorter or longer than the internodes, smooth or scabridulous, tightly imbricate, yellowish-brown, basal sheaths laterally compressed, keeled, not becoming spirally coiled when old;

ligules 3-12 mm, firm and brown basally, membranous distally, acute;

blades 10-45 cm long, 1.6-5 mm wide, tightly folded, scabrous abaxially, hirsute adaxially.

Panicles

3-15 cm long, 1-1.6 cm wide, contracted, often interrupted below;

branches 0.4-5.5 cm, appressed or diverging up to 30° from the rachises, spikelet-bearing to the base;

pedicels 0.1-4 mm, scabrous to hirsute;

disarticulation above the glumes.

18-40 cm long, 1.5-7 cm wide, loosely contracted to open, not dense;

branches 1-10 cm, ascending or diverging up to 60° from the rachises, stiff, naked basally;

pedicels 2-8 mm, hirtellous.

Spikelets

1.8-2.8 mm, borne singly.

3-4.2 mm, yellowish to purplish.

Glumes

unequal, shorter than the florets, thin and membranous throughout, unawned;

lower glumes lacking or rudimentary, veinless, rounded and often erose;

upper glumes 0.1-0.3 mm, veinless;

lemmas 1.8-2.8 mm, oblong-elliptic, mostly scabrous, calluses hairy, hairs to 0.8 mm, veins greenish, lower 1/4 of the midveins with a few appressed hairs, apices acute to acuminate, awned, awns 1.5-5 mm, straight;

paleas 1.8-2.8 mm, oblong-elliptic, acute to acuminate;

anthers 0.2-0.5 mm, yellow.

subequal, 2-3 mm, exceeded by the florets, scabridulous or smooth, 1-veined, acute or obtuse, unawned;

lemmas 3-4.2 mm, lanceolate, glabrous or appressed-pubescent on the lower 1/4 of the margins, apices acute to obtuse, usually bifid and awned, teeth to 0.3 mm, awns 0.5-4 mm, straight;

paleas 3-4.2 mm, lanceolate, glabrous, acute to acuminate;

anthers 1-1.8 mm, purplish.

Caryopses

1-1.4 mm, fusiform, brownish.

not seen.

2n

= 40, 42.

= 24.

Muhlenbergia schreberi

Muhlenbergia ×involuta

Distribution
from FNA
AL; AR; AZ; CA; CO; CT; DC; DE; FL; GA; IA; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; MS; NC; NE; NH; NJ; NY; OH; OK; PA; RI; SC; SD; TN; TX; UT; VA; VT; WI; WV; ON
[WildflowerSearch map]
[BONAP county map]
from FNA
TX
[BONAP county map]
Discussion

Muhlenbergia schreberi grows in moist to dry woods and prairies on rocky slopes, in ravines, and along sandy riverbanks, at elevations of 60-1600 m. It is also common in disturbed sites near cultivated fields, pastures, and roads at these elevations. Its geographic range includes central, but not northern, Mexico. Records from the western United States probably reflect receent introductions. The species is considered a noxious, invasive weed in California.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Muhlenbergia ×involuta grows on rocky, calcareous slopes in openings and along canyons, at elevations of 150-500 m. It has only been found growing naturally in Texas, but it is also available commercially as an ornamental. Swallen (1932) suggested that M. reverchonii and M. lindheimeri were its parents, but M. rigida seems to be another plausible possibility.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 25, p. 162. FNA vol. 25, p. 187.
Parent taxa Poaceae > subfam. Chloridoideae > tribe Cynodonteae > Muhlenbergia Poaceae > subfam. Chloridoideae > tribe Cynodonteae > Muhlenbergia
Sibling taxa
M. andina, M. appressa, M. arenacea, M. arenicola, M. arizonica, M. arsenei, M. asperifolia, M. brevis, M. bushii, M. californica, M. capillaris, M. crispiseta, M. curtifolia, M. cuspidata, M. depauperata, M. diversiglumis, M. dubia, M. dumosa, M. elongata, M. eludens, M. emersleyi, M. expansa, M. filiculmis, M. filiformis, M. fragilis, M. frondosa, M. glabrifloris, M. glauca, M. glomerata, M. jonesii, M. lindheimeri, M. longiligula, M. mexicana, M. microsperma, M. minutissima, M. montana, M. palmeri, M. pauciflora, M. pectinata, M. peruviana, M. polycaulis, M. porteri, M. pungens, M. racemosa, M. ramulosa, M. repens, M. reverchonii, M. richardsonis, M. rigens, M. rigida, M. sericea, M. setifolia, M. sinuosa, M. sobolifera, M. spiciformis, M. straminea, M. sylvatica, M. tenuiflora, M. tenuifolia, M. texana, M. thurberi, M. torreyana, M. torreyi, M. uniflora, M. utilis, M. villiflora, M. wrightii, M. ×curtisetosa, M. ×involuta
M. andina, M. appressa, M. arenacea, M. arenicola, M. arizonica, M. arsenei, M. asperifolia, M. brevis, M. bushii, M. californica, M. capillaris, M. crispiseta, M. curtifolia, M. cuspidata, M. depauperata, M. diversiglumis, M. dubia, M. dumosa, M. elongata, M. eludens, M. emersleyi, M. expansa, M. filiculmis, M. filiformis, M. fragilis, M. frondosa, M. glabrifloris, M. glauca, M. glomerata, M. jonesii, M. lindheimeri, M. longiligula, M. mexicana, M. microsperma, M. minutissima, M. montana, M. palmeri, M. pauciflora, M. pectinata, M. peruviana, M. polycaulis, M. porteri, M. pungens, M. racemosa, M. ramulosa, M. repens, M. reverchonii, M. richardsonis, M. rigens, M. rigida, M. schreberi, M. sericea, M. setifolia, M. sinuosa, M. sobolifera, M. spiciformis, M. straminea, M. sylvatica, M. tenuiflora, M. tenuifolia, M. texana, M. thurberi, M. torreyana, M. torreyi, M. uniflora, M. utilis, M. villiflora, M. wrightii, M. ×curtisetosa
Synonyms M. palustris, M. diffusa
Name authority J.E Gmel. Swallen
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