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nimblewell, nimblewill, nimblewill muhly

forest muhly, muhlenbergie des bois, woodland muhly

Habit Plants perennial (appearing annual); usually cespitose, not rhizomatous, sometimes stoloniferous. Plants perennial; rhizomatous.
Culms

10-45(70) cm, geniculate, often rooting at the lower nodes, glabrous or puberulent below the nodes;

internodes often smooth, shiny, glabrous.

40-110 cm tall, 1-2 mm thick, erect;

internodes puberulent for most of their length, strigose below the nodes.

Sheaths

shorter than the internodes, glabrous for most of their length, margins shortly (0.3-1.2 mm) pubescent distally, not becoming spirally coiled when old;

ligules 0.2-0.5 mm, truncate, erose, ciliate;

blades (1)3-10 cm long, 1-4.5 mm wide, flat, smooth or scabridulous.

glabrous and smooth for most of their length, scabridulous distally, margins hyaline;

ligules 1-2.5 mm, membranous, truncate, lacerate-ciliolate;

blades 5-18 cm long, 3-7 mm wide, flat, scabrous to scabridulous, occasionally smooth.

Panicles

3-15 cm long, 1-1.6 cm wide, contracted, often interrupted below;

branches 0.4-5.5 cm, appressed or diverging up to 30° from the rachises, spikelet-bearing to the base;

pedicels 0.1-4 mm, scabrous to hirsute;

disarticulation above the glumes.

terminal and axillary, 6-21 cm long, 0.2-1 cm wide, narrow, not dense;

axillary panicles usually exserted at maturity;

branches 0.8-6 cm, ascending to closely appressed;

pedicels 0.8-3.5 mm, strigose.

Spikelets

1.8-2.8 mm, borne singly.

2.2-3.7 mm.

Glumes

unequal, shorter than the florets, thin and membranous throughout, unawned;

lower glumes lacking or rudimentary, veinless, rounded and often erose;

upper glumes 0.1-0.3 mm, veinless;

lemmas 1.8-2.8 mm, oblong-elliptic, mostly scabrous, calluses hairy, hairs to 0.8 mm, veins greenish, lower 1/4 of the midveins with a few appressed hairs, apices acute to acuminate, awned, awns 1.5-5 mm, straight;

paleas 1.8-2.8 mm, oblong-elliptic, acute to acuminate;

anthers 0.2-0.5 mm, yellow.

subequal, 1.8-3 mm, nearly as long as the lemmas, 1-veined, tapering from near the base, apices scabridulous, acuminate, unawned or awned, awns to 1 mm;

lemmas 2.2-3.7 mm, lanceolate to narrowly lanceolate, hairy on the calluses, lower 1/2 of the midveins, and margins, hairs 0.2-0.5 mm, apices scabridulous, acuminate, awned, awns 5-18 mm, purplish;

paleas 2-3.5 mm, lanceolate, proximal 1/2 shortly pilose, apices scabridulous, acuminate;

anthers 0.4-0.8 mm, yellow.

Caryopses

1-1.4 mm, fusiform, brownish.

1.4-2 mm, fusiform, brown.

2n

= 40, 42.

= 40.

Muhlenbergia schreberi

Muhlenbergia sylvatica

Distribution
from FNA
AL; AR; AZ; CA; CO; CT; DC; DE; FL; GA; IA; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; MS; NC; NE; NH; NJ; NY; OH; OK; PA; RI; SC; SD; TN; TX; UT; VA; VT; WI; WV; ON
[WildflowerSearch map]
[BONAP county map]
from FNA
AL; AR; AZ; CT; DC; DE; GA; IA; IL; IN; KS; KY; MA; MD; ME; MI; MN; MO; MS; NC; NE; NH; NJ; NY; OH; OK; PA; RI; SC; SD; TN; TX; VA; VT; WI; WV; ON; QC
[WildflowerSearch map]
[BONAP county map]
Discussion

Muhlenbergia schreberi grows in moist to dry woods and prairies on rocky slopes, in ravines, and along sandy riverbanks, at elevations of 60-1600 m. It is also common in disturbed sites near cultivated fields, pastures, and roads at these elevations. Its geographic range includes central, but not northern, Mexico. Records from the western United States probably reflect receent introductions. The species is considered a noxious, invasive weed in California.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Muhlenbergia sylvatica grows in upland forests, along creeks and hollows, on rocky ledges derived from sandstone, shale, or calcareous parent materials, moist prairies, and swamps, at elevations from 30-1500 m. It is restricted to the Flora region, its primary range being southeastern Canada and the midwestern and eastern United States. Reports from British Columbia were based on a misidentification (Douglas et al. 2002). The record from Arizona is based on the report in Kearney and Peebles (1951) of a collection made by Toumey at Grapevine Creek in the Grand Canyon.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 25, p. 162. FNA vol. 25, p. 160.
Parent taxa Poaceae > subfam. Chloridoideae > tribe Cynodonteae > Muhlenbergia Poaceae > subfam. Chloridoideae > tribe Cynodonteae > Muhlenbergia
Sibling taxa
M. andina, M. appressa, M. arenacea, M. arenicola, M. arizonica, M. arsenei, M. asperifolia, M. brevis, M. bushii, M. californica, M. capillaris, M. crispiseta, M. curtifolia, M. cuspidata, M. depauperata, M. diversiglumis, M. dubia, M. dumosa, M. elongata, M. eludens, M. emersleyi, M. expansa, M. filiculmis, M. filiformis, M. fragilis, M. frondosa, M. glabrifloris, M. glauca, M. glomerata, M. jonesii, M. lindheimeri, M. longiligula, M. mexicana, M. microsperma, M. minutissima, M. montana, M. palmeri, M. pauciflora, M. pectinata, M. peruviana, M. polycaulis, M. porteri, M. pungens, M. racemosa, M. ramulosa, M. repens, M. reverchonii, M. richardsonis, M. rigens, M. rigida, M. sericea, M. setifolia, M. sinuosa, M. sobolifera, M. spiciformis, M. straminea, M. sylvatica, M. tenuiflora, M. tenuifolia, M. texana, M. thurberi, M. torreyana, M. torreyi, M. uniflora, M. utilis, M. villiflora, M. wrightii, M. ×curtisetosa, M. ×involuta
M. andina, M. appressa, M. arenacea, M. arenicola, M. arizonica, M. arsenei, M. asperifolia, M. brevis, M. bushii, M. californica, M. capillaris, M. crispiseta, M. curtifolia, M. cuspidata, M. depauperata, M. diversiglumis, M. dubia, M. dumosa, M. elongata, M. eludens, M. emersleyi, M. expansa, M. filiculmis, M. filiformis, M. fragilis, M. frondosa, M. glabrifloris, M. glauca, M. glomerata, M. jonesii, M. lindheimeri, M. longiligula, M. mexicana, M. microsperma, M. minutissima, M. montana, M. palmeri, M. pauciflora, M. pectinata, M. peruviana, M. polycaulis, M. porteri, M. pungens, M. racemosa, M. ramulosa, M. repens, M. reverchonii, M. richardsonis, M. rigens, M. rigida, M. schreberi, M. sericea, M. setifolia, M. sinuosa, M. sobolifera, M. spiciformis, M. straminea, M. tenuiflora, M. tenuifolia, M. texana, M. thurberi, M. torreyana, M. torreyi, M. uniflora, M. utilis, M. villiflora, M. wrightii, M. ×curtisetosa, M. ×involuta
Synonyms M. palustris, M. diffusa M. sylvatica var. robusta, M. sylvatica forma attenuata
Name authority J.E Gmel. (Torn) Torr. ex A. Gray
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