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nimblewell, nimblewill, nimblewill muhly

seep muhly

Habit Plants perennial (appearing annual); usually cespitose, not rhizomatous, sometimes stoloniferous. Plants perennial; cespitose, not rhizomatous, typical bunch-grasses in appearance.
Culms

10-45(70) cm, geniculate, often rooting at the lower nodes, glabrous or puberulent below the nodes;

internodes often smooth, shiny, glabrous.

40-80 cm, stiffly erect from the base, not conspicuously branched;

internodes glabrous, sometimes puberulent below the nodes.

Sheaths

shorter than the internodes, glabrous for most of their length, margins shortly (0.3-1.2 mm) pubescent distally, not becoming spirally coiled when old;

ligules 0.2-0.5 mm, truncate, erose, ciliate;

blades (1)3-10 cm long, 1-4.5 mm wide, flat, smooth or scabridulous.

shorter than the internodes, smooth or scabridulous, basal sheaths rounded, not becoming spirally coiled when old;

ligules (2)4-7(9) mm, firmer near the base, obtuse, lacerate;

blades 8-35 cm long, 1-2 mm wide, flat or involute, filiform, glabrous abaxially, densely hirtellous adaxially.

Panicles

3-15 cm long, 1-1.6 cm wide, contracted, often interrupted below;

branches 0.4-5.5 cm, appressed or diverging up to 30° from the rachises, spikelet-bearing to the base;

pedicels 0.1-4 mm, scabrous to hirsute;

disarticulation above the glumes.

10-20(30) cm long, 4-15 cm wide, about as wide as long, loosely contracted to open but not diffuse;

primary branches 1.4-10 cm, capillary, diverging up to 80° from the rachises, scabridulous, naked basally, lower branches with 5-15(20) spikelets;

pedicels 3-25 mm, longer than the spikelets, capillary, stiff, or flexuous.

Spikelets

1.8-2.8 mm, borne singly.

3.5-5 mm, stramineous or brownish to purplish.

Glumes

unequal, shorter than the florets, thin and membranous throughout, unawned;

lower glumes lacking or rudimentary, veinless, rounded and often erose;

upper glumes 0.1-0.3 mm, veinless;

lemmas 1.8-2.8 mm, oblong-elliptic, mostly scabrous, calluses hairy, hairs to 0.8 mm, veins greenish, lower 1/4 of the midveins with a few appressed hairs, apices acute to acuminate, awned, awns 1.5-5 mm, straight;

paleas 1.8-2.8 mm, oblong-elliptic, acute to acuminate;

anthers 0.2-0.5 mm, yellow.

subequal, 1-3 mm, shorter than the florets, hyaline, mostly sparsely hirtellous, apices glabrous, 1-veined (sometimes faintly so), acute, occasionally mucronate, mucros shorter than 0.7 mm;

lemmas 3.5-5 mm, narrowly lanceolate, calluses hairy, hairs to 0.5 mm, lemma bodies glabrous and smooth, apices scabridulous, acuminate, awned, awns 0.5-4(6) mm, clearly demarcated from the lemma bodies;

paleas 3.5-5 mm, narrowly lanceolate, mostly glabrous;

anthers 1.1-2 mm, yellowish to purplish.

Caryopses

1-1.4 mm, fusiform, brownish.

2-2.4 mm, fusiform, brownish.

2n

= 40, 42.

= 20.

Muhlenbergia schreberi

Muhlenbergia reverchonii

Distribution
from FNA
AL; AR; AZ; CA; CO; CT; DC; DE; FL; GA; IA; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; MS; NC; NE; NH; NJ; NY; OH; OK; PA; RI; SC; SD; TN; TX; UT; VA; VT; WI; WV; ON
[WildflowerSearch map]
[BONAP county map]
from FNA
OK; TX
[BONAP county map]
Discussion

Muhlenbergia schreberi grows in moist to dry woods and prairies on rocky slopes, in ravines, and along sandy riverbanks, at elevations of 60-1600 m. It is also common in disturbed sites near cultivated fields, pastures, and roads at these elevations. Its geographic range includes central, but not northern, Mexico. Records from the western United States probably reflect receent introductions. The species is considered a noxious, invasive weed in California.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Muhlenbergia reverchonii grows on calcareous rocky slopes, flats, and limestone rock outcrops, at elevations of 150-650 m. It is restricted to Oklahoma and Texas. Muhlenbergia reverchonii resembles M. capillaris and M. setifolia in many respects, but differs from the former in its smooth and shiny lemmas, and from the latter in its wider panicles, spreading panicle branches, acute glumes, and more shortly awned lemmas.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 25, p. 162. FNA vol. 25, p. 190.
Parent taxa Poaceae > subfam. Chloridoideae > tribe Cynodonteae > Muhlenbergia Poaceae > subfam. Chloridoideae > tribe Cynodonteae > Muhlenbergia
Sibling taxa
M. andina, M. appressa, M. arenacea, M. arenicola, M. arizonica, M. arsenei, M. asperifolia, M. brevis, M. bushii, M. californica, M. capillaris, M. crispiseta, M. curtifolia, M. cuspidata, M. depauperata, M. diversiglumis, M. dubia, M. dumosa, M. elongata, M. eludens, M. emersleyi, M. expansa, M. filiculmis, M. filiformis, M. fragilis, M. frondosa, M. glabrifloris, M. glauca, M. glomerata, M. jonesii, M. lindheimeri, M. longiligula, M. mexicana, M. microsperma, M. minutissima, M. montana, M. palmeri, M. pauciflora, M. pectinata, M. peruviana, M. polycaulis, M. porteri, M. pungens, M. racemosa, M. ramulosa, M. repens, M. reverchonii, M. richardsonis, M. rigens, M. rigida, M. sericea, M. setifolia, M. sinuosa, M. sobolifera, M. spiciformis, M. straminea, M. sylvatica, M. tenuiflora, M. tenuifolia, M. texana, M. thurberi, M. torreyana, M. torreyi, M. uniflora, M. utilis, M. villiflora, M. wrightii, M. ×curtisetosa, M. ×involuta
M. andina, M. appressa, M. arenacea, M. arenicola, M. arizonica, M. arsenei, M. asperifolia, M. brevis, M. bushii, M. californica, M. capillaris, M. crispiseta, M. curtifolia, M. cuspidata, M. depauperata, M. diversiglumis, M. dubia, M. dumosa, M. elongata, M. eludens, M. emersleyi, M. expansa, M. filiculmis, M. filiformis, M. fragilis, M. frondosa, M. glabrifloris, M. glauca, M. glomerata, M. jonesii, M. lindheimeri, M. longiligula, M. mexicana, M. microsperma, M. minutissima, M. montana, M. palmeri, M. pauciflora, M. pectinata, M. peruviana, M. polycaulis, M. porteri, M. pungens, M. racemosa, M. ramulosa, M. repens, M. richardsonis, M. rigens, M. rigida, M. schreberi, M. sericea, M. setifolia, M. sinuosa, M. sobolifera, M. spiciformis, M. straminea, M. sylvatica, M. tenuiflora, M. tenuifolia, M. texana, M. thurberi, M. torreyana, M. torreyi, M. uniflora, M. utilis, M. villiflora, M. wrightii, M. ×curtisetosa, M. ×involuta
Synonyms M. palustris, M. diffusa
Name authority J.E Gmel. Vasey & Scribn.
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