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nimblewell, nimblewill, nimblewill muhly

cliff muhly

Habit Plants perennial (appearing annual); usually cespitose, not rhizomatous, sometimes stoloniferous. Plants perennial; loosely cespitose, sometimes shortly rhizomatous.
Culms

10-45(70) cm, geniculate, often rooting at the lower nodes, glabrous or puberulent below the nodes;

internodes often smooth, shiny, glabrous.

15-40(50) cm tall, 0.8-1.5 mm thick, erect or decumbent;

internodes glabrous or strigulose for most of their length, including below the nodes.

Sheaths

shorter than the internodes, glabrous for most of their length, margins shortly (0.3-1.2 mm) pubescent distally, not becoming spirally coiled when old;

ligules 0.2-0.5 mm, truncate, erose, ciliate;

blades (1)3-10 cm long, 1-4.5 mm wide, flat, smooth or scabridulous.

usually shorter than the internodes, smooth or scabridulous, without necrotic spots, not becoming spirally coiled when old;

ligules 0.5-2.5 mm, membranous, obtuse to acute, without lateral lobes;

blades 3-10 mm long, 0.5-2 mm wide, flat or involute (occasionally folded), smooth or scabridulous abaxially, hirsute or scabridulous adaxially, without necrotic spots.

Panicles

3-15 cm long, 1-1.6 cm wide, contracted, often interrupted below;

branches 0.4-5.5 cm, appressed or diverging up to 30° from the rachises, spikelet-bearing to the base;

pedicels 0.1-4 mm, scabrous to hirsute;

disarticulation above the glumes.

2-12 cm long, 0.3-1.8 cm wide, contracted, interrupted below;

primary branches 0.5-4 cm, appressed or diverging up to 30° from the rachises, spikelet-bearing to the base;

pedicels 0.1-1.5 mm, scabrous.

Spikelets

1.8-2.8 mm, borne singly.

2.5-4 mm, plump near the middle.

Glumes

unequal, shorter than the florets, thin and membranous throughout, unawned;

lower glumes lacking or rudimentary, veinless, rounded and often erose;

upper glumes 0.1-0.3 mm, veinless;

lemmas 1.8-2.8 mm, oblong-elliptic, mostly scabrous, calluses hairy, hairs to 0.8 mm, veins greenish, lower 1/4 of the midveins with a few appressed hairs, apices acute to acuminate, awned, awns 1.5-5 mm, straight;

paleas 1.8-2.8 mm, oblong-elliptic, acute to acuminate;

anthers 0.2-0.5 mm, yellow.

subequal, (1)1.5-2.6 mm, exceeded by the florets, 1-veined, acute or acuminate, unawned or awned, awns to 1.4 mm;

lemmas 2-3.5 mm, elliptic, widest near the middle, appressed-pubescent on the lower 1/2 - 2/3 of the midveins and margins, hairs to 0.5 mm, apices scabridulous, acuminate, awned, awns 10-20(25) mm, flexuous;

paleas 2-3.5 mm, elliptic, intercostal region appressed-pubescent on the basal 1/2, apices acuminate;

anthers 1.5-2 mm, orange.

Caryopses

1-1.4 mm, fusiform, brownish.

1.5-2 mm, fusiform, brownish.

2n

= 40, 42.

= 20, 40.

Muhlenbergia schreberi

Muhlenbergia polycaulis

Distribution
from FNA
AL; AR; AZ; CA; CO; CT; DC; DE; FL; GA; IA; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; MS; NC; NE; NH; NJ; NY; OH; OK; PA; RI; SC; SD; TN; TX; UT; VA; VT; WI; WV; ON
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; NM; TX; UT
[BONAP county map]
Discussion

Muhlenbergia schreberi grows in moist to dry woods and prairies on rocky slopes, in ravines, and along sandy riverbanks, at elevations of 60-1600 m. It is also common in disturbed sites near cultivated fields, pastures, and roads at these elevations. Its geographic range includes central, but not northern, Mexico. Records from the western United States probably reflect receent introductions. The species is considered a noxious, invasive weed in California.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Muhlenbergia polycaulis grows in open vegetation on steep rocky slopes, canyon walls, cliffs, table rocks, and volcanic rock outcrops, at elevations of 1200-2400 m. Its range extends from the southwestern United States to central Mexico. It differs from M. glauca, with which it may be confused, in its longer lemma awns, shorter rhizomes, and loosely tufted habit.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 25, p. 162. FNA vol. 25, p. 167.
Parent taxa Poaceae > subfam. Chloridoideae > tribe Cynodonteae > Muhlenbergia Poaceae > subfam. Chloridoideae > tribe Cynodonteae > Muhlenbergia
Sibling taxa
M. andina, M. appressa, M. arenacea, M. arenicola, M. arizonica, M. arsenei, M. asperifolia, M. brevis, M. bushii, M. californica, M. capillaris, M. crispiseta, M. curtifolia, M. cuspidata, M. depauperata, M. diversiglumis, M. dubia, M. dumosa, M. elongata, M. eludens, M. emersleyi, M. expansa, M. filiculmis, M. filiformis, M. fragilis, M. frondosa, M. glabrifloris, M. glauca, M. glomerata, M. jonesii, M. lindheimeri, M. longiligula, M. mexicana, M. microsperma, M. minutissima, M. montana, M. palmeri, M. pauciflora, M. pectinata, M. peruviana, M. polycaulis, M. porteri, M. pungens, M. racemosa, M. ramulosa, M. repens, M. reverchonii, M. richardsonis, M. rigens, M. rigida, M. sericea, M. setifolia, M. sinuosa, M. sobolifera, M. spiciformis, M. straminea, M. sylvatica, M. tenuiflora, M. tenuifolia, M. texana, M. thurberi, M. torreyana, M. torreyi, M. uniflora, M. utilis, M. villiflora, M. wrightii, M. ×curtisetosa, M. ×involuta
M. andina, M. appressa, M. arenacea, M. arenicola, M. arizonica, M. arsenei, M. asperifolia, M. brevis, M. bushii, M. californica, M. capillaris, M. crispiseta, M. curtifolia, M. cuspidata, M. depauperata, M. diversiglumis, M. dubia, M. dumosa, M. elongata, M. eludens, M. emersleyi, M. expansa, M. filiculmis, M. filiformis, M. fragilis, M. frondosa, M. glabrifloris, M. glauca, M. glomerata, M. jonesii, M. lindheimeri, M. longiligula, M. mexicana, M. microsperma, M. minutissima, M. montana, M. palmeri, M. pauciflora, M. pectinata, M. peruviana, M. porteri, M. pungens, M. racemosa, M. ramulosa, M. repens, M. reverchonii, M. richardsonis, M. rigens, M. rigida, M. schreberi, M. sericea, M. setifolia, M. sinuosa, M. sobolifera, M. spiciformis, M. straminea, M. sylvatica, M. tenuiflora, M. tenuifolia, M. texana, M. thurberi, M. torreyana, M. torreyi, M. uniflora, M. utilis, M. villiflora, M. wrightii, M. ×curtisetosa, M. ×involuta
Synonyms M. palustris, M. diffusa
Name authority J.E Gmel. Scribn.
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