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nimblewell, nimblewill, nimblewill muhly

Peruvian muhly

Habit Plants perennial (appearing annual); usually cespitose, not rhizomatous, sometimes stoloniferous. Plants annual; tufted.
Culms

10-45(70) cm, geniculate, often rooting at the lower nodes, glabrous or puberulent below the nodes;

internodes often smooth, shiny, glabrous.

3-27 cm, erect, glabrous.

Sheaths

shorter than the internodes, glabrous for most of their length, margins shortly (0.3-1.2 mm) pubescent distally, not becoming spirally coiled when old;

ligules 0.2-0.5 mm, truncate, erose, ciliate;

blades (1)3-10 cm long, 1-4.5 mm wide, flat, smooth or scabridulous.

usually longer than the internodes, smooth or scabridulous;

ligules 1.5-3 mm, membra nous, acute;

blades 1-5 cm long, 0.6-1.5 mm wide, flat to involute, smooth or scabridulous abaxially, sometimes shortly pubescent adaxially.

Panicles

3-15 cm long, 1-1.6 cm wide, contracted, often interrupted below;

branches 0.4-5.5 cm, appressed or diverging up to 30° from the rachises, spikelet-bearing to the base;

pedicels 0.1-4 mm, scabrous to hirsute;

disarticulation above the glumes.

2-8 cm long, 0.3-3.4 cm wide, contracted or open;

primary branches 1-5 cm, diverging up to 80° from the rachises;

pedicels 0.4-5 mm, smooth or scabrous.

Spikelets

1.8-2.8 mm, borne singly.

1.4-4.2 mm.

Glumes

unequal, shorter than the florets, thin and membranous throughout, unawned;

lower glumes lacking or rudimentary, veinless, rounded and often erose;

upper glumes 0.1-0.3 mm, veinless;

lemmas 1.8-2.8 mm, oblong-elliptic, mostly scabrous, calluses hairy, hairs to 0.8 mm, veins greenish, lower 1/4 of the midveins with a few appressed hairs, apices acute to acuminate, awned, awns 1.5-5 mm, straight;

paleas 1.8-2.8 mm, oblong-elliptic, acute to acuminate;

anthers 0.2-0.5 mm, yellow.

smooth or scabridulous;

lower glumes 0.8-2.8 mm, narrow to broadly lanceolate, 1-veined, acute, often awn-tipped;

upper glumes 0.9-3 mm, wider than the lower glumes, lanceolate, (1)2-3-veined, truncate to acute, 2- or 3-toothed;

lemmas 1.4-4.2 mm, widest near the base, purplish mottled with dark green, hairy on the calluses and lower M of the lemma bodies, hairs to 0.5 mm, apices acuminate, usually bifid and awned from between the teeth, teeth to 0.5 mm, awns 3-10 mm, flexuous, purplish;

paleas 1.3-3.8 mm, narrowly lanceolate, acuminate to subacute;

anthers 0.5-1 mm, purplish to yellowish.

Caryopses

1-1.4 mm, fusiform, brownish.

1-1.6 mm, fusiform, brownish.

2n

= 40, 42.

= 30.

Muhlenbergia schreberi

Muhlenbergia peruviana

Distribution
from FNA
AL; AR; AZ; CA; CO; CT; DC; DE; FL; GA; IA; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; MS; NC; NE; NH; NJ; NY; OH; OK; PA; RI; SC; SD; TN; TX; UT; VA; VT; WI; WV; ON
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; NM
[BONAP county map]
Discussion

Muhlenbergia schreberi grows in moist to dry woods and prairies on rocky slopes, in ravines, and along sandy riverbanks, at elevations of 60-1600 m. It is also common in disturbed sites near cultivated fields, pastures, and roads at these elevations. Its geographic range includes central, but not northern, Mexico. Records from the western United States probably reflect receent introductions. The species is considered a noxious, invasive weed in California.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Muhlenbergia peruviana grows in open gravelly flats, meadows, rock outcrops, sandy washes, gravelly drainages, rocky slopes, disturbed road cuts, and volcanic flats, in yellow pine forest associations, at elevations of 2000-4600 m. Its primary distribution is to the south of the Flora region, extending from the southwestern United States through Mexico to Ecuador, Peru, Bolivia, and Argentina.

As treated here, Muhlenbergia peruviana includes what are sometimes identified as M. pulcherrima Scribn. ex Beal, M. pusilla Steud., and M. peruviana s. s. There are, however, numerous intermediates among the three extremes represented by these names.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 25, p. 162. FNA vol. 25, p. 185.
Parent taxa Poaceae > subfam. Chloridoideae > tribe Cynodonteae > Muhlenbergia Poaceae > subfam. Chloridoideae > tribe Cynodonteae > Muhlenbergia
Sibling taxa
M. andina, M. appressa, M. arenacea, M. arenicola, M. arizonica, M. arsenei, M. asperifolia, M. brevis, M. bushii, M. californica, M. capillaris, M. crispiseta, M. curtifolia, M. cuspidata, M. depauperata, M. diversiglumis, M. dubia, M. dumosa, M. elongata, M. eludens, M. emersleyi, M. expansa, M. filiculmis, M. filiformis, M. fragilis, M. frondosa, M. glabrifloris, M. glauca, M. glomerata, M. jonesii, M. lindheimeri, M. longiligula, M. mexicana, M. microsperma, M. minutissima, M. montana, M. palmeri, M. pauciflora, M. pectinata, M. peruviana, M. polycaulis, M. porteri, M. pungens, M. racemosa, M. ramulosa, M. repens, M. reverchonii, M. richardsonis, M. rigens, M. rigida, M. sericea, M. setifolia, M. sinuosa, M. sobolifera, M. spiciformis, M. straminea, M. sylvatica, M. tenuiflora, M. tenuifolia, M. texana, M. thurberi, M. torreyana, M. torreyi, M. uniflora, M. utilis, M. villiflora, M. wrightii, M. ×curtisetosa, M. ×involuta
M. andina, M. appressa, M. arenacea, M. arenicola, M. arizonica, M. arsenei, M. asperifolia, M. brevis, M. bushii, M. californica, M. capillaris, M. crispiseta, M. curtifolia, M. cuspidata, M. depauperata, M. diversiglumis, M. dubia, M. dumosa, M. elongata, M. eludens, M. emersleyi, M. expansa, M. filiculmis, M. filiformis, M. fragilis, M. frondosa, M. glabrifloris, M. glauca, M. glomerata, M. jonesii, M. lindheimeri, M. longiligula, M. mexicana, M. microsperma, M. minutissima, M. montana, M. palmeri, M. pauciflora, M. pectinata, M. polycaulis, M. porteri, M. pungens, M. racemosa, M. ramulosa, M. repens, M. reverchonii, M. richardsonis, M. rigens, M. rigida, M. schreberi, M. sericea, M. setifolia, M. sinuosa, M. sobolifera, M. spiciformis, M. straminea, M. sylvatica, M. tenuiflora, M. tenuifolia, M. texana, M. thurberi, M. torreyana, M. torreyi, M. uniflora, M. utilis, M. villiflora, M. wrightii, M. ×curtisetosa, M. ×involuta
Synonyms M. palustris, M. diffusa M. pulcherrima
Name authority J.E Gmel. (P. Beauv.) Steud.
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