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nimblewell, nimblewill, nimblewill muhly

mixedglume muhly

Habit Plants perennial (appearing annual); usually cespitose, not rhizomatous, sometimes stoloniferous. Plants annual; sprawling.
Culms

10-45(70) cm, geniculate, often rooting at the lower nodes, glabrous or puberulent below the nodes;

internodes often smooth, shiny, glabrous.

16-50 cm, decumbent, rooting at the lower nodes;

nodes retrorsely pilose;

internodes smooth or scabridulous.

Sheaths

shorter than the internodes, glabrous for most of their length, margins shortly (0.3-1.2 mm) pubescent distally, not becoming spirally coiled when old;

ligules 0.2-0.5 mm, truncate, erose, ciliate;

blades (1)3-10 cm long, 1-4.5 mm wide, flat, smooth or scabridulous.

1.5-8.5 cm, sparsely or densely pilose, hairs to 3 mm, papillose-based;

ligules 0.5-0.8 mm, membranous, truncate, erose;

blades 2-6 cm long, 1.5-4 mm wide, flat, bases distinctly narrowed to the junction with the sheath, surfaces scabridulous and sparsely pilose, hairs papillose-based.

Panicles

3-15 cm long, 1-1.6 cm wide, contracted, often interrupted below;

branches 0.4-5.5 cm, appressed or diverging up to 30° from the rachises, spikelet-bearing to the base;

pedicels 0.1-4 mm, scabrous to hirsute;

disarticulation above the glumes.

6-10.5 cm long, 2.0-4.5 cm wide, secund, open;

primary branches 0.8-3.5 cm, secund, spreading at right angles or somewhat reflexed, with 2-5 spikelets;

secondary branches not developed;

pedicels 1-5 mm, scabrous or shortly pilose, hairs papillose-based;

disarticulation at the base of the primary branches.

Spikelets

1.8-2.8 mm, borne singly.

4-8 mm, dimorphic with respect to the glumes, proximal spikelets on each branch almost sessile.

Glumes

unequal, shorter than the florets, thin and membranous throughout, unawned;

lower glumes lacking or rudimentary, veinless, rounded and often erose;

upper glumes 0.1-0.3 mm, veinless;

lemmas 1.8-2.8 mm, oblong-elliptic, mostly scabrous, calluses hairy, hairs to 0.8 mm, veins greenish, lower 1/4 of the midveins with a few appressed hairs, apices acute to acuminate, awned, awns 1.5-5 mm, straight;

paleas 1.8-2.8 mm, oblong-elliptic, acute to acuminate;

anthers 0.2-0.5 mm, yellow.

of proximal spikelets on each branch subequal, 0.2-0.7 mm, orbicular, truncate, often erose, unawned;

glumes of distal spikelets on each branch markedly unequal;

lower glumes to 8 mm, 1-veined, acute, usually awned, awns 0.5-3 mm;

upper glumes orbicular, acute, sometimes awn-tipped;

lemmas 4.0-7.6 mm, linear to broadly lanceolate, light greenish, smooth or scabrous, usually with greenish veins, apices acuminate, awned, awns 6-19 mm, usually straight, scabrous;

paleas 3.7-6.8 mm, narrowly lanceolate, coarsely papillate or almost smooth, acuminate;

anthers 0.4-0.8 mm, yellowish.

Caryopses

1-1.4 mm, fusiform, brownish.

1.8-3 mm, oblong-ovoid, flattened, brownish.

2n

= 40, 42.

= 20.

Muhlenbergia schreberi

Muhlenbergia diversiglumis

Distribution
from FNA
AL; AR; AZ; CA; CO; CT; DC; DE; FL; GA; IA; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; MS; NC; NE; NH; NJ; NY; OH; OK; PA; RI; SC; SD; TN; TX; UT; VA; VT; WI; WV; ON
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Discussion

Muhlenbergia schreberi grows in moist to dry woods and prairies on rocky slopes, in ravines, and along sandy riverbanks, at elevations of 60-1600 m. It is also common in disturbed sites near cultivated fields, pastures, and roads at these elevations. Its geographic range includes central, but not northern, Mexico. Records from the western United States probably reflect receent introductions. The species is considered a noxious, invasive weed in California.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Muhlenbergia diversiglumis has been collected from Galveston County, Texas. The species is native from Mexico to Peru and Venezuela, where it grows on moist cliffs, along water courses, sandy slopes, and road cuts, primarily in moist shaded environments of broadleaf evergreen forests and pine-oak forests, at elevations of 600-2500 m. The collection from Texas may represent a recent introduction.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 25, p. 162. FNA vol. 25, p. 164.
Parent taxa Poaceae > subfam. Chloridoideae > tribe Cynodonteae > Muhlenbergia Poaceae > subfam. Chloridoideae > tribe Cynodonteae > Muhlenbergia
Sibling taxa
M. andina, M. appressa, M. arenacea, M. arenicola, M. arizonica, M. arsenei, M. asperifolia, M. brevis, M. bushii, M. californica, M. capillaris, M. crispiseta, M. curtifolia, M. cuspidata, M. depauperata, M. diversiglumis, M. dubia, M. dumosa, M. elongata, M. eludens, M. emersleyi, M. expansa, M. filiculmis, M. filiformis, M. fragilis, M. frondosa, M. glabrifloris, M. glauca, M. glomerata, M. jonesii, M. lindheimeri, M. longiligula, M. mexicana, M. microsperma, M. minutissima, M. montana, M. palmeri, M. pauciflora, M. pectinata, M. peruviana, M. polycaulis, M. porteri, M. pungens, M. racemosa, M. ramulosa, M. repens, M. reverchonii, M. richardsonis, M. rigens, M. rigida, M. sericea, M. setifolia, M. sinuosa, M. sobolifera, M. spiciformis, M. straminea, M. sylvatica, M. tenuiflora, M. tenuifolia, M. texana, M. thurberi, M. torreyana, M. torreyi, M. uniflora, M. utilis, M. villiflora, M. wrightii, M. ×curtisetosa, M. ×involuta
M. andina, M. appressa, M. arenacea, M. arenicola, M. arizonica, M. arsenei, M. asperifolia, M. brevis, M. bushii, M. californica, M. capillaris, M. crispiseta, M. curtifolia, M. cuspidata, M. depauperata, M. dubia, M. dumosa, M. elongata, M. eludens, M. emersleyi, M. expansa, M. filiculmis, M. filiformis, M. fragilis, M. frondosa, M. glabrifloris, M. glauca, M. glomerata, M. jonesii, M. lindheimeri, M. longiligula, M. mexicana, M. microsperma, M. minutissima, M. montana, M. palmeri, M. pauciflora, M. pectinata, M. peruviana, M. polycaulis, M. porteri, M. pungens, M. racemosa, M. ramulosa, M. repens, M. reverchonii, M. richardsonis, M. rigens, M. rigida, M. schreberi, M. sericea, M. setifolia, M. sinuosa, M. sobolifera, M. spiciformis, M. straminea, M. sylvatica, M. tenuiflora, M. tenuifolia, M. texana, M. thurberi, M. torreyana, M. torreyi, M. uniflora, M. utilis, M. villiflora, M. wrightii, M. ×curtisetosa, M. ×involuta
Synonyms M. palustris, M. diffusa
Name authority J.E Gmel. Trin.
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