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mat muhly, Matted muhly, muhlenbergie de Richardson, Richardson's muhly, soft-leaf muhly

seep muhly

Habit Plants perennial; rhizomatous, not cespitose, often mat-forming. Plants perennial; cespitose, not rhizomatous, typical bunch-grasses in appearance.
Culms

5-30 cm tall, 0.4-1 mm thick, decumbent, geniculate, or erect;

internodes usually nodulose (occasionally smooth) for most of their length, puberulent or nodulose below the nodes.

40-80 cm, stiffly erect from the base, not conspicuously branched;

internodes glabrous, sometimes puberulent below the nodes.

Sheaths

shorter or longer than the internodes, glabrous;

ligules 0.8-3 mm, membranous, acute to truncate, erose;

blades 0.4-6.5 cm long, 0.5-4.2 mm wide, flat or involute, straight or arcuate-spreading, glabrous abaxially, hirtellous adaxially.

shorter than the internodes, smooth or scabridulous, basal sheaths rounded, not becoming spirally coiled when old;

ligules (2)4-7(9) mm, firmer near the base, obtuse, lacerate;

blades 8-35 cm long, 1-2 mm wide, flat or involute, filiform, glabrous abaxially, densely hirtellous adaxially.

Panicles

1-15 cm long, 0.1-1.7 cm wide, exserted, narrow or spikelike, rachises usually concealed by the branches;

primary branches 0.4-5 cm, usually closely appressed at maturity, rarely diverging up to 20° from the rachises;

pedicels 0.2-2 mm, setulose.

10-20(30) cm long, 4-15 cm wide, about as wide as long, loosely contracted to open but not diffuse;

primary branches 1.4-10 cm, capillary, diverging up to 80° from the rachises, scabridulous, naked basally, lower branches with 5-15(20) spikelets;

pedicels 3-25 mm, longer than the spikelets, capillary, stiff, or flexuous.

Spikelets

1.7-3.1 mm, occasionally with 2 florets.

3.5-5 mm, stramineous or brownish to purplish.

Glumes

subequal, 0.6-2 mm, 1/3– 1/2 as long as the lemmas, green, 1(2)-veined, acute, sometimes mucronate, mucros less than 0.2 mm;

lemmas 1.7-2.6(3.1) mm, lanceolate, dark greenish, plumbeous, or mottled, glabrous, apices scabridulous, acute to acuminate, sometimes mucronate, mucros to 0.5 mm;

paleas 1.2-2.4(2.9) mm, lanceolate, acute;

anthers 0.9-1.6 mm, yellow to purplish.

subequal, 1-3 mm, shorter than the florets, hyaline, mostly sparsely hirtellous, apices glabrous, 1-veined (sometimes faintly so), acute, occasionally mucronate, mucros shorter than 0.7 mm;

lemmas 3.5-5 mm, narrowly lanceolate, calluses hairy, hairs to 0.5 mm, lemma bodies glabrous and smooth, apices scabridulous, acuminate, awned, awns 0.5-4(6) mm, clearly demarcated from the lemma bodies;

paleas 3.5-5 mm, narrowly lanceolate, mostly glabrous;

anthers 1.1-2 mm, yellowish to purplish.

Caryopses

0.9-1.6 mm, narrowly ellipsoid, brown.

2-2.4 mm, fusiform, brownish.

2n

= 40.

= 20.

Muhlenbergia richardsonis

Muhlenbergia reverchonii

Distribution
from FNA
AK; AZ; CA; CO; ID; ME; MI; MN; MT; ND; NE; NM; NV; OR; SD; UT; WA; WI; WY; AB; BC; MB; NB; NT; ON; QC; SK; YT
[WildflowerSearch map]
[BONAP county map]
from FNA
OK; TX
[BONAP county map]
Discussion

Muhlenbergia richardsonis grows in open sites in alkaline meadows, prairies, sandy arroyo bottoms, talus slopes, rocky flats and the shores of rivers, at elevations of 60-3300 m. It is the most widespread species of Muhlenbergia in the Flora region, extending from the Yukon Territory to Quebec in the north and to northern Baja California, Mexico, in the south. Morden and Hatch (1996) reported that it also grows in Alaska, but no voucher specimen has been located.

Muhlenbergia richardsonis is often confused with M. cuspidata, which differs in lacking rhizomes and having shorter ligules, and sometimes with M. filiformis, which differs in being a weak annual with glabrous internodes and obtuse, erose glumes.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Muhlenbergia reverchonii grows on calcareous rocky slopes, flats, and limestone rock outcrops, at elevations of 150-650 m. It is restricted to Oklahoma and Texas. Muhlenbergia reverchonii resembles M. capillaris and M. setifolia in many respects, but differs from the former in its smooth and shiny lemmas, and from the latter in its wider panicles, spreading panicle branches, acute glumes, and more shortly awned lemmas.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 25, p. 177. FNA vol. 25, p. 190.
Parent taxa Poaceae > subfam. Chloridoideae > tribe Cynodonteae > Muhlenbergia Poaceae > subfam. Chloridoideae > tribe Cynodonteae > Muhlenbergia
Sibling taxa
M. andina, M. appressa, M. arenacea, M. arenicola, M. arizonica, M. arsenei, M. asperifolia, M. brevis, M. bushii, M. californica, M. capillaris, M. crispiseta, M. curtifolia, M. cuspidata, M. depauperata, M. diversiglumis, M. dubia, M. dumosa, M. elongata, M. eludens, M. emersleyi, M. expansa, M. filiculmis, M. filiformis, M. fragilis, M. frondosa, M. glabrifloris, M. glauca, M. glomerata, M. jonesii, M. lindheimeri, M. longiligula, M. mexicana, M. microsperma, M. minutissima, M. montana, M. palmeri, M. pauciflora, M. pectinata, M. peruviana, M. polycaulis, M. porteri, M. pungens, M. racemosa, M. ramulosa, M. repens, M. reverchonii, M. rigens, M. rigida, M. schreberi, M. sericea, M. setifolia, M. sinuosa, M. sobolifera, M. spiciformis, M. straminea, M. sylvatica, M. tenuiflora, M. tenuifolia, M. texana, M. thurberi, M. torreyana, M. torreyi, M. uniflora, M. utilis, M. villiflora, M. wrightii, M. ×curtisetosa, M. ×involuta
M. andina, M. appressa, M. arenacea, M. arenicola, M. arizonica, M. arsenei, M. asperifolia, M. brevis, M. bushii, M. californica, M. capillaris, M. crispiseta, M. curtifolia, M. cuspidata, M. depauperata, M. diversiglumis, M. dubia, M. dumosa, M. elongata, M. eludens, M. emersleyi, M. expansa, M. filiculmis, M. filiformis, M. fragilis, M. frondosa, M. glabrifloris, M. glauca, M. glomerata, M. jonesii, M. lindheimeri, M. longiligula, M. mexicana, M. microsperma, M. minutissima, M. montana, M. palmeri, M. pauciflora, M. pectinata, M. peruviana, M. polycaulis, M. porteri, M. pungens, M. racemosa, M. ramulosa, M. repens, M. richardsonis, M. rigens, M. rigida, M. schreberi, M. sericea, M. setifolia, M. sinuosa, M. sobolifera, M. spiciformis, M. straminea, M. sylvatica, M. tenuiflora, M. tenuifolia, M. texana, M. thurberi, M. torreyana, M. torreyi, M. uniflora, M. utilis, M. villiflora, M. wrightii, M. ×curtisetosa, M. ×involuta
Synonyms Sporobolus depauperatus, M. squarrosa
Name authority (Trin.) Rydb. Vasey & Scribn.
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