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mat muhly, Matted muhly, muhlenbergie de Richardson, Richardson's muhly, soft-leaf muhly

cliff muhly

Habit Plants perennial; rhizomatous, not cespitose, often mat-forming. Plants perennial; loosely cespitose, sometimes shortly rhizomatous.
Culms

5-30 cm tall, 0.4-1 mm thick, decumbent, geniculate, or erect;

internodes usually nodulose (occasionally smooth) for most of their length, puberulent or nodulose below the nodes.

15-40(50) cm tall, 0.8-1.5 mm thick, erect or decumbent;

internodes glabrous or strigulose for most of their length, including below the nodes.

Sheaths

shorter or longer than the internodes, glabrous;

ligules 0.8-3 mm, membranous, acute to truncate, erose;

blades 0.4-6.5 cm long, 0.5-4.2 mm wide, flat or involute, straight or arcuate-spreading, glabrous abaxially, hirtellous adaxially.

usually shorter than the internodes, smooth or scabridulous, without necrotic spots, not becoming spirally coiled when old;

ligules 0.5-2.5 mm, membranous, obtuse to acute, without lateral lobes;

blades 3-10 mm long, 0.5-2 mm wide, flat or involute (occasionally folded), smooth or scabridulous abaxially, hirsute or scabridulous adaxially, without necrotic spots.

Panicles

1-15 cm long, 0.1-1.7 cm wide, exserted, narrow or spikelike, rachises usually concealed by the branches;

primary branches 0.4-5 cm, usually closely appressed at maturity, rarely diverging up to 20° from the rachises;

pedicels 0.2-2 mm, setulose.

2-12 cm long, 0.3-1.8 cm wide, contracted, interrupted below;

primary branches 0.5-4 cm, appressed or diverging up to 30° from the rachises, spikelet-bearing to the base;

pedicels 0.1-1.5 mm, scabrous.

Spikelets

1.7-3.1 mm, occasionally with 2 florets.

2.5-4 mm, plump near the middle.

Glumes

subequal, 0.6-2 mm, 1/3– 1/2 as long as the lemmas, green, 1(2)-veined, acute, sometimes mucronate, mucros less than 0.2 mm;

lemmas 1.7-2.6(3.1) mm, lanceolate, dark greenish, plumbeous, or mottled, glabrous, apices scabridulous, acute to acuminate, sometimes mucronate, mucros to 0.5 mm;

paleas 1.2-2.4(2.9) mm, lanceolate, acute;

anthers 0.9-1.6 mm, yellow to purplish.

subequal, (1)1.5-2.6 mm, exceeded by the florets, 1-veined, acute or acuminate, unawned or awned, awns to 1.4 mm;

lemmas 2-3.5 mm, elliptic, widest near the middle, appressed-pubescent on the lower 1/2 - 2/3 of the midveins and margins, hairs to 0.5 mm, apices scabridulous, acuminate, awned, awns 10-20(25) mm, flexuous;

paleas 2-3.5 mm, elliptic, intercostal region appressed-pubescent on the basal 1/2, apices acuminate;

anthers 1.5-2 mm, orange.

Caryopses

0.9-1.6 mm, narrowly ellipsoid, brown.

1.5-2 mm, fusiform, brownish.

2n

= 40.

= 20, 40.

Muhlenbergia richardsonis

Muhlenbergia polycaulis

Distribution
from FNA
AK; AZ; CA; CO; ID; ME; MI; MN; MT; ND; NE; NM; NV; OR; SD; UT; WA; WI; WY; AB; BC; MB; NB; NT; ON; QC; SK; YT
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; NM; TX; UT
[BONAP county map]
Discussion

Muhlenbergia richardsonis grows in open sites in alkaline meadows, prairies, sandy arroyo bottoms, talus slopes, rocky flats and the shores of rivers, at elevations of 60-3300 m. It is the most widespread species of Muhlenbergia in the Flora region, extending from the Yukon Territory to Quebec in the north and to northern Baja California, Mexico, in the south. Morden and Hatch (1996) reported that it also grows in Alaska, but no voucher specimen has been located.

Muhlenbergia richardsonis is often confused with M. cuspidata, which differs in lacking rhizomes and having shorter ligules, and sometimes with M. filiformis, which differs in being a weak annual with glabrous internodes and obtuse, erose glumes.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Muhlenbergia polycaulis grows in open vegetation on steep rocky slopes, canyon walls, cliffs, table rocks, and volcanic rock outcrops, at elevations of 1200-2400 m. Its range extends from the southwestern United States to central Mexico. It differs from M. glauca, with which it may be confused, in its longer lemma awns, shorter rhizomes, and loosely tufted habit.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 25, p. 177. FNA vol. 25, p. 167.
Parent taxa Poaceae > subfam. Chloridoideae > tribe Cynodonteae > Muhlenbergia Poaceae > subfam. Chloridoideae > tribe Cynodonteae > Muhlenbergia
Sibling taxa
M. andina, M. appressa, M. arenacea, M. arenicola, M. arizonica, M. arsenei, M. asperifolia, M. brevis, M. bushii, M. californica, M. capillaris, M. crispiseta, M. curtifolia, M. cuspidata, M. depauperata, M. diversiglumis, M. dubia, M. dumosa, M. elongata, M. eludens, M. emersleyi, M. expansa, M. filiculmis, M. filiformis, M. fragilis, M. frondosa, M. glabrifloris, M. glauca, M. glomerata, M. jonesii, M. lindheimeri, M. longiligula, M. mexicana, M. microsperma, M. minutissima, M. montana, M. palmeri, M. pauciflora, M. pectinata, M. peruviana, M. polycaulis, M. porteri, M. pungens, M. racemosa, M. ramulosa, M. repens, M. reverchonii, M. rigens, M. rigida, M. schreberi, M. sericea, M. setifolia, M. sinuosa, M. sobolifera, M. spiciformis, M. straminea, M. sylvatica, M. tenuiflora, M. tenuifolia, M. texana, M. thurberi, M. torreyana, M. torreyi, M. uniflora, M. utilis, M. villiflora, M. wrightii, M. ×curtisetosa, M. ×involuta
M. andina, M. appressa, M. arenacea, M. arenicola, M. arizonica, M. arsenei, M. asperifolia, M. brevis, M. bushii, M. californica, M. capillaris, M. crispiseta, M. curtifolia, M. cuspidata, M. depauperata, M. diversiglumis, M. dubia, M. dumosa, M. elongata, M. eludens, M. emersleyi, M. expansa, M. filiculmis, M. filiformis, M. fragilis, M. frondosa, M. glabrifloris, M. glauca, M. glomerata, M. jonesii, M. lindheimeri, M. longiligula, M. mexicana, M. microsperma, M. minutissima, M. montana, M. palmeri, M. pauciflora, M. pectinata, M. peruviana, M. porteri, M. pungens, M. racemosa, M. ramulosa, M. repens, M. reverchonii, M. richardsonis, M. rigens, M. rigida, M. schreberi, M. sericea, M. setifolia, M. sinuosa, M. sobolifera, M. spiciformis, M. straminea, M. sylvatica, M. tenuiflora, M. tenuifolia, M. texana, M. thurberi, M. torreyana, M. torreyi, M. uniflora, M. utilis, M. villiflora, M. wrightii, M. ×curtisetosa, M. ×involuta
Synonyms Sporobolus depauperatus, M. squarrosa
Name authority (Trin.) Rydb. Scribn.
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