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mat muhly, Matted muhly, muhlenbergie de Richardson, Richardson's muhly, soft-leaf muhly

littleseed muhly

Habit Plants perennial; rhizomatous, not cespitose, often mat-forming. Plants annual, sometimes appearing as short-lived perennials; tufted.
Culms

5-30 cm tall, 0.4-1 mm thick, decumbent, geniculate, or erect;

internodes usually nodulose (occasionally smooth) for most of their length, puberulent or nodulose below the nodes.

10-80 cm, often geniculate at the base, much branched near the base;

internodes mostly scabridulous or smooth, always scabridulous below the nodes.

Sheaths

shorter or longer than the internodes, glabrous;

ligules 0.8-3 mm, membranous, acute to truncate, erose;

blades 0.4-6.5 cm long, 0.5-4.2 mm wide, flat or involute, straight or arcuate-spreading, glabrous abaxially, hirtellous adaxially.

often shorter than the internodes, glabrous, smooth or scabridulous;

ligules 1-2 mm, membranous to hyaline, truncate to obtuse;

blades 3-8.5(10) cm long, 1-2.5 mm wide, flat or loosely involute, scabrous abaxially, strigulose adaxially.

Panicles

1-15 cm long, 0.1-1.7 cm wide, exserted, narrow or spikelike, rachises usually concealed by the branches;

primary branches 0.4-5 cm, usually closely appressed at maturity, rarely diverging up to 20° from the rachises;

pedicels 0.2-2 mm, setulose.

6.5-13.5 cm long, 1-6.5 cm wide, not dense, often purplish;

branches 1.6-4 cm, ascending or diverging up to 80° from the rachises, spikelet-bearing to the base;

pedicels 2-6 mm, appressed to divaricate, antrorsely scabrous;

disarticulation above the glumes.

Spikelets

1.7-3.1 mm, occasionally with 2 florets.

2.5-5.5 mm, borne singly.

Glumes

subequal, 0.6-2 mm, 1/3– 1/2 as long as the lemmas, green, 1(2)-veined, acute, sometimes mucronate, mucros less than 0.2 mm;

lemmas 1.7-2.6(3.1) mm, lanceolate, dark greenish, plumbeous, or mottled, glabrous, apices scabridulous, acute to acuminate, sometimes mucronate, mucros to 0.5 mm;

paleas 1.2-2.4(2.9) mm, lanceolate, acute;

anthers 0.9-1.6 mm, yellow to purplish.

0.4-1.3 mm, exceeded by the florets, 1-veined, obtuse, often minutely erose;

lower glumes 0.4-1 mm;

upper glumes 0.6-1.3 mm;

lemmas 2.5-3.8(5.3) mm, narrowly lanceolate, mostly smooth, scabridulous distally, hairy on the calluses, lower 1/2 of the margins, and midveins, hairs 0.2-0.5 mm, apices acuminate, awned, awns 10-30 mm, straight to flexuous;

paleas 2.2-4.8 mm, narrowly lanceolate, acuminate;

anthers 0.3-1.2 mm, purplish.

Caryopses

0.9-1.6 mm, narrowly ellipsoid, brown.

1.7-2.5 mm, fusiform, reddish-brown.

Cleistogamous

panicles with 1-3 spikelets present in the axils of the lower leaves.

2n

= 40.

= 20, 40, 60.

Muhlenbergia richardsonis

Muhlenbergia microsperma

Distribution
from FNA
AK; AZ; CA; CO; ID; ME; MI; MN; MT; ND; NE; NM; NV; OR; SD; UT; WA; WI; WY; AB; BC; MB; NB; NT; ON; QC; SK; YT
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; CA; NV; UT; HI
[WildflowerSearch map]
[BONAP county map]
Discussion

Muhlenbergia richardsonis grows in open sites in alkaline meadows, prairies, sandy arroyo bottoms, talus slopes, rocky flats and the shores of rivers, at elevations of 60-3300 m. It is the most widespread species of Muhlenbergia in the Flora region, extending from the Yukon Territory to Quebec in the north and to northern Baja California, Mexico, in the south. Morden and Hatch (1996) reported that it also grows in Alaska, but no voucher specimen has been located.

Muhlenbergia richardsonis is often confused with M. cuspidata, which differs in lacking rhizomes and having shorter ligules, and sometimes with M. filiformis, which differs in being a weak annual with glabrous internodes and obtuse, erose glumes.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Muhlenbergia microsperma grows on sandy slopes, drainages, cliffs, rock outcrops, and disturbed roadsides, at elevations of 0-2400 m. It is usually found in creosote scrub, thorn-scrub forest, sarcocaulescent desert, and oak-pinyon woodland associations. Its range extends from the southwestern United States through Central America to Peru and Venezuela. Morphological variation among and within its populations is marked.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 25, p. 177. FNA vol. 25, p. 162.
Parent taxa Poaceae > subfam. Chloridoideae > tribe Cynodonteae > Muhlenbergia Poaceae > subfam. Chloridoideae > tribe Cynodonteae > Muhlenbergia
Sibling taxa
M. andina, M. appressa, M. arenacea, M. arenicola, M. arizonica, M. arsenei, M. asperifolia, M. brevis, M. bushii, M. californica, M. capillaris, M. crispiseta, M. curtifolia, M. cuspidata, M. depauperata, M. diversiglumis, M. dubia, M. dumosa, M. elongata, M. eludens, M. emersleyi, M. expansa, M. filiculmis, M. filiformis, M. fragilis, M. frondosa, M. glabrifloris, M. glauca, M. glomerata, M. jonesii, M. lindheimeri, M. longiligula, M. mexicana, M. microsperma, M. minutissima, M. montana, M. palmeri, M. pauciflora, M. pectinata, M. peruviana, M. polycaulis, M. porteri, M. pungens, M. racemosa, M. ramulosa, M. repens, M. reverchonii, M. rigens, M. rigida, M. schreberi, M. sericea, M. setifolia, M. sinuosa, M. sobolifera, M. spiciformis, M. straminea, M. sylvatica, M. tenuiflora, M. tenuifolia, M. texana, M. thurberi, M. torreyana, M. torreyi, M. uniflora, M. utilis, M. villiflora, M. wrightii, M. ×curtisetosa, M. ×involuta
M. andina, M. appressa, M. arenacea, M. arenicola, M. arizonica, M. arsenei, M. asperifolia, M. brevis, M. bushii, M. californica, M. capillaris, M. crispiseta, M. curtifolia, M. cuspidata, M. depauperata, M. diversiglumis, M. dubia, M. dumosa, M. elongata, M. eludens, M. emersleyi, M. expansa, M. filiculmis, M. filiformis, M. fragilis, M. frondosa, M. glabrifloris, M. glauca, M. glomerata, M. jonesii, M. lindheimeri, M. longiligula, M. mexicana, M. minutissima, M. montana, M. palmeri, M. pauciflora, M. pectinata, M. peruviana, M. polycaulis, M. porteri, M. pungens, M. racemosa, M. ramulosa, M. repens, M. reverchonii, M. richardsonis, M. rigens, M. rigida, M. schreberi, M. sericea, M. setifolia, M. sinuosa, M. sobolifera, M. spiciformis, M. straminea, M. sylvatica, M. tenuiflora, M. tenuifolia, M. texana, M. thurberi, M. torreyana, M. torreyi, M. uniflora, M. utilis, M. villiflora, M. wrightii, M. ×curtisetosa, M. ×involuta
Synonyms Sporobolus depauperatus, M. squarrosa
Name authority (Trin.) Rydb. (DC.) Trin.
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