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mat muhly, Matted muhly, muhlenbergie de Richardson, Richardson's muhly, soft-leaf muhly

bristly muhly, marsh muhly, muhlenbergie agglomeree, spike muhly

Habit Plants perennial; rhizomatous, not cespitose, often mat-forming. Plants perennial; rhizomatous, not cespitose.
Culms

5-30 cm tall, 0.4-1 mm thick, decumbent, geniculate, or erect;

internodes usually nodulose (occasionally smooth) for most of their length, puberulent or nodulose below the nodes.

30-120 cm tall, 0.8-2.5 mm thick, erect, seldom branched above the base;

internodes dull, mostly puberulent (sometimes sparsely so), terete, rarely keeled, strigose immediately below the nodes.

Sheaths

shorter or longer than the internodes, glabrous;

ligules 0.8-3 mm, membranous, acute to truncate, erose;

blades 0.4-6.5 cm long, 0.5-4.2 mm wide, flat or involute, straight or arcuate-spreading, glabrous abaxially, hirtellous adaxially.

scabridulous, slightly keeled;

ligules 0.2-0.6 mm, membranous, truncate, lacerate-ciliolate;

blades 2-15 cm long, 2-6 mm wide, flat, usually scabrous or scabridulous, occasionally smooth.

Panicles

1-15 cm long, 0.1-1.7 cm wide, exserted, narrow or spikelike, rachises usually concealed by the branches;

primary branches 0.4-5 cm, usually closely appressed at maturity, rarely diverging up to 20° from the rachises;

pedicels 0.2-2 mm, setulose.

1.5-12 cm long, 0.3-1.8 cm wide, lobed, dense;

primary branches 0.2-2.5 cm, appressed;

pedicels absent or to 1 mm, strigose.

Spikelets

1.7-3.1 mm, occasionally with 2 florets.

3-8 mm.

Glumes

subequal, 0.6-2 mm, 1/3– 1/2 as long as the lemmas, green, 1(2)-veined, acute, sometimes mucronate, mucros less than 0.2 mm;

lemmas 1.7-2.6(3.1) mm, lanceolate, dark greenish, plumbeous, or mottled, glabrous, apices scabridulous, acute to acuminate, sometimes mucronate, mucros to 0.5 mm;

paleas 1.2-2.4(2.9) mm, lanceolate, acute;

anthers 0.9-1.6 mm, yellow to purplish.

subequal, 3-8 mm (including the awn), about 1.3-2 times longer than the lemmas, smooth or scabridulous distally, 1-veined, acuminate, awned, awns to 5 mm;

lemmas 1.9-3.1 mm, lanceolate, pubescent on the calluses, mid-veins, and margins, hairs to 1.2 mm, apices scabridulous, acuminate, unawned or awned, awns to 1 mm;

paleas 1.9-3.1 mm, lanceolate, loosely pilose between the veins, apices acuminate;

anthers 0.8-1.5 mm, yellowish.

Caryopses

0.9-1.6 mm, narrowly ellipsoid, brown.

1-1.6 mm, fusiform, brown.

2n

= 40.

= 20.

Muhlenbergia richardsonis

Muhlenbergia glomerata

Distribution
from FNA
AK; AZ; CA; CO; ID; ME; MI; MN; MT; ND; NE; NM; NV; OR; SD; UT; WA; WI; WY; AB; BC; MB; NB; NT; ON; QC; SK; YT
[WildflowerSearch map]
[BONAP county map]
from FNA
CO; CT; DC; IA; ID; IL; IN; KY; MA; MD; ME; MI; MN; MT; NC; ND; NE; NH; NJ; NV; NY; OH; OR; PA; RI; SD; UT; VA; VT; WA; WI; WV; WY; AB; BC; LB; MB; NB; NS; NT; ON; PE; QC; SK; YT
[WildflowerSearch map]
[BONAP county map]
Discussion

Muhlenbergia richardsonis grows in open sites in alkaline meadows, prairies, sandy arroyo bottoms, talus slopes, rocky flats and the shores of rivers, at elevations of 60-3300 m. It is the most widespread species of Muhlenbergia in the Flora region, extending from the Yukon Territory to Quebec in the north and to northern Baja California, Mexico, in the south. Morden and Hatch (1996) reported that it also grows in Alaska, but no voucher specimen has been located.

Muhlenbergia richardsonis is often confused with M. cuspidata, which differs in lacking rhizomes and having shorter ligules, and sometimes with M. filiformis, which differs in being a weak annual with glabrous internodes and obtuse, erose glumes.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Muhlenbergia glomerata grows in meadows, marshes, bogs, alkaline fens, lake margins, stream banks, beside irrigation ditches and hot springs, and on gravelly slopes, in many different plant communities, at elevations of 30-2300 m. It is most common in southern Canada and the northeastern United States, but grows sporadically throughout the western United States. It is not known from Mexico.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 25, p. 177. FNA vol. 25, p. 154.
Parent taxa Poaceae > subfam. Chloridoideae > tribe Cynodonteae > Muhlenbergia Poaceae > subfam. Chloridoideae > tribe Cynodonteae > Muhlenbergia
Sibling taxa
M. andina, M. appressa, M. arenacea, M. arenicola, M. arizonica, M. arsenei, M. asperifolia, M. brevis, M. bushii, M. californica, M. capillaris, M. crispiseta, M. curtifolia, M. cuspidata, M. depauperata, M. diversiglumis, M. dubia, M. dumosa, M. elongata, M. eludens, M. emersleyi, M. expansa, M. filiculmis, M. filiformis, M. fragilis, M. frondosa, M. glabrifloris, M. glauca, M. glomerata, M. jonesii, M. lindheimeri, M. longiligula, M. mexicana, M. microsperma, M. minutissima, M. montana, M. palmeri, M. pauciflora, M. pectinata, M. peruviana, M. polycaulis, M. porteri, M. pungens, M. racemosa, M. ramulosa, M. repens, M. reverchonii, M. rigens, M. rigida, M. schreberi, M. sericea, M. setifolia, M. sinuosa, M. sobolifera, M. spiciformis, M. straminea, M. sylvatica, M. tenuiflora, M. tenuifolia, M. texana, M. thurberi, M. torreyana, M. torreyi, M. uniflora, M. utilis, M. villiflora, M. wrightii, M. ×curtisetosa, M. ×involuta
M. andina, M. appressa, M. arenacea, M. arenicola, M. arizonica, M. arsenei, M. asperifolia, M. brevis, M. bushii, M. californica, M. capillaris, M. crispiseta, M. curtifolia, M. cuspidata, M. depauperata, M. diversiglumis, M. dubia, M. dumosa, M. elongata, M. eludens, M. emersleyi, M. expansa, M. filiculmis, M. filiformis, M. fragilis, M. frondosa, M. glabrifloris, M. glauca, M. jonesii, M. lindheimeri, M. longiligula, M. mexicana, M. microsperma, M. minutissima, M. montana, M. palmeri, M. pauciflora, M. pectinata, M. peruviana, M. polycaulis, M. porteri, M. pungens, M. racemosa, M. ramulosa, M. repens, M. reverchonii, M. richardsonis, M. rigens, M. rigida, M. schreberi, M. sericea, M. setifolia, M. sinuosa, M. sobolifera, M. spiciformis, M. straminea, M. sylvatica, M. tenuiflora, M. tenuifolia, M. texana, M. thurberi, M. torreyana, M. torreyi, M. uniflora, M. utilis, M. villiflora, M. wrightii, M. ×curtisetosa, M. ×involuta
Synonyms Sporobolus depauperatus, M. squarrosa M. racemosa var. cinnoides, M. glomerata var. cinnoides
Name authority (Trin.) Rydb. (Willd.) Trin.
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