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mat muhly, Matted muhly, muhlenbergie de Richardson, Richardson's muhly, soft-leaf muhly

pull-up muhly, slender muhly

Habit Plants perennial; rhizomatous, not cespitose, often mat-forming. Plants annual (often appearing perennial), tufted.
Culms

5-30 cm tall, 0.4-1 mm thick, decumbent, geniculate, or erect;

internodes usually nodulose (occasionally smooth) for most of their length, puberulent or nodulose below the nodes.

(3)5-20(35) cm, erect or geniculate, often rooting at the lower nodes;

internodes glabrous.

Sheaths

shorter or longer than the internodes, glabrous;

ligules 0.8-3 mm, membranous, acute to truncate, erose;

blades 0.4-6.5 cm long, 0.5-4.2 mm wide, flat or involute, straight or arcuate-spreading, glabrous abaxially, hirtellous adaxially.

shorter or longer than the internodes, glabrous, smooth or scabridulous;

ligules 1-3.5 mm, hyaline to membranous, rounded to acute;

blades 1-4(6) cm long, 0.6-1.6 mm wide, flat or involute, smooth or scabridulous abaxially, scabrous or pubescent adaxially.

Panicles

1-15 cm long, 0.1-1.7 cm wide, exserted, narrow or spikelike, rachises usually concealed by the branches;

primary branches 0.4-5 cm, usually closely appressed at maturity, rarely diverging up to 20° from the rachises;

pedicels 0.2-2 mm, setulose.

1.6-6 cm long, 0.2-0.5 cm wide, spikelike, interrupted near the base, long-exserted;

primary branches 0.9-1.2 cm, closely appressed at maturity;

pedicels 1-3 mm, scabrous.

Spikelets

1.7-3.1 mm, occasionally with 2 florets.

1.5-3.2 mm.

Glumes

subequal, 0.6-2 mm, 1/3– 1/2 as long as the lemmas, green, 1(2)-veined, acute, sometimes mucronate, mucros less than 0.2 mm;

lemmas 1.7-2.6(3.1) mm, lanceolate, dark greenish, plumbeous, or mottled, glabrous, apices scabridulous, acute to acuminate, sometimes mucronate, mucros to 0.5 mm;

paleas 1.2-2.4(2.9) mm, lanceolate, acute;

anthers 0.9-1.6 mm, yellow to purplish.

greenish-gray, glabrous, 1-veined, rounded to subacute;

lower glumes 0.6-1.4 mm;

upper glumes 0.7-1.7 mm;

lemmas (1.5)1.8-2.5(3.2) mm, lanceolate, dark greenish, appressed-pubescent on the margins and midveins, hairs shorter than 0.3 mm, apices scabridulous, acute to acuminate, unawned, sometimes mucronate, mucros shorter than 1 mm;

paleas 1.6-2.6(3.1) mm, lanceolate, scabridulous distally;

anthers 0.5-1.2 mm, purplish.

Caryopses

0.9-1.6 mm, narrowly ellipsoid, brown.

0.9-1.5 mm, fusiform, reddish-brown.

2n

= 40.

= 18.

Muhlenbergia richardsonis

Muhlenbergia filiformis

Distribution
from FNA
AK; AZ; CA; CO; ID; ME; MI; MN; MT; ND; NE; NM; NV; OR; SD; UT; WA; WI; WY; AB; BC; MB; NB; NT; ON; QC; SK; YT
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; CA; CO; ID; KS; MT; ND; NE; NM; NV; OR; SD; UT; WA; WY; BC
[WildflowerSearch map]
[BONAP county map]
Discussion

Muhlenbergia richardsonis grows in open sites in alkaline meadows, prairies, sandy arroyo bottoms, talus slopes, rocky flats and the shores of rivers, at elevations of 60-3300 m. It is the most widespread species of Muhlenbergia in the Flora region, extending from the Yukon Territory to Quebec in the north and to northern Baja California, Mexico, in the south. Morden and Hatch (1996) reported that it also grows in Alaska, but no voucher specimen has been located.

Muhlenbergia richardsonis is often confused with M. cuspidata, which differs in lacking rhizomes and having shorter ligules, and sometimes with M. filiformis, which differs in being a weak annual with glabrous internodes and obtuse, erose glumes.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Muhlenbergia filiformis grows in open, moist meadows, on gravelly lake shores, along stream banks, and in moist humus near thermal springs, at elevations of 1060-3050 m. It is usually associated with yellow pine forests, but also grows in many other plant communities. Its range extends into northern Mexico.

Muhlenbergia filiformis resembles M. richardsonis, but differs in having glabrous internodes and subacute apices. Large, robust specimens have been referred to M. simplex Scribn. or M. filiformis var. fortis E.H. Kelso but, until there is more evidence to the contrary, it seems best to treat such plants as representing an extreme of the variation within M. filiformis.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 25, p. 177. FNA vol. 25, p. 179.
Parent taxa Poaceae > subfam. Chloridoideae > tribe Cynodonteae > Muhlenbergia Poaceae > subfam. Chloridoideae > tribe Cynodonteae > Muhlenbergia
Sibling taxa
M. andina, M. appressa, M. arenacea, M. arenicola, M. arizonica, M. arsenei, M. asperifolia, M. brevis, M. bushii, M. californica, M. capillaris, M. crispiseta, M. curtifolia, M. cuspidata, M. depauperata, M. diversiglumis, M. dubia, M. dumosa, M. elongata, M. eludens, M. emersleyi, M. expansa, M. filiculmis, M. filiformis, M. fragilis, M. frondosa, M. glabrifloris, M. glauca, M. glomerata, M. jonesii, M. lindheimeri, M. longiligula, M. mexicana, M. microsperma, M. minutissima, M. montana, M. palmeri, M. pauciflora, M. pectinata, M. peruviana, M. polycaulis, M. porteri, M. pungens, M. racemosa, M. ramulosa, M. repens, M. reverchonii, M. rigens, M. rigida, M. schreberi, M. sericea, M. setifolia, M. sinuosa, M. sobolifera, M. spiciformis, M. straminea, M. sylvatica, M. tenuiflora, M. tenuifolia, M. texana, M. thurberi, M. torreyana, M. torreyi, M. uniflora, M. utilis, M. villiflora, M. wrightii, M. ×curtisetosa, M. ×involuta
M. andina, M. appressa, M. arenacea, M. arenicola, M. arizonica, M. arsenei, M. asperifolia, M. brevis, M. bushii, M. californica, M. capillaris, M. crispiseta, M. curtifolia, M. cuspidata, M. depauperata, M. diversiglumis, M. dubia, M. dumosa, M. elongata, M. eludens, M. emersleyi, M. expansa, M. filiculmis, M. fragilis, M. frondosa, M. glabrifloris, M. glauca, M. glomerata, M. jonesii, M. lindheimeri, M. longiligula, M. mexicana, M. microsperma, M. minutissima, M. montana, M. palmeri, M. pauciflora, M. pectinata, M. peruviana, M. polycaulis, M. porteri, M. pungens, M. racemosa, M. ramulosa, M. repens, M. reverchonii, M. richardsonis, M. rigens, M. rigida, M. schreberi, M. sericea, M. setifolia, M. sinuosa, M. sobolifera, M. spiciformis, M. straminea, M. sylvatica, M. tenuiflora, M. tenuifolia, M. texana, M. thurberi, M. torreyana, M. torreyi, M. uniflora, M. utilis, M. villiflora, M. wrightii, M. ×curtisetosa, M. ×involuta
Synonyms Sporobolus depauperatus, M. squarrosa Sporobolus gracillimus
Name authority (Trin.) Rydb. (Thurb. ex S. Watson) Rydb.
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