Muhlenbergia richardsonis |
Muhlenbergia diversiglumis |
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mat muhly, Matted muhly, muhlenbergie de Richardson, Richardson's muhly, soft-leaf muhly |
mixedglume muhly |
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Habit | Plants perennial; rhizomatous, not cespitose, often mat-forming. | Plants annual; sprawling. |
Culms | 5-30 cm tall, 0.4-1 mm thick, decumbent, geniculate, or erect; internodes usually nodulose (occasionally smooth) for most of their length, puberulent or nodulose below the nodes. |
16-50 cm, decumbent, rooting at the lower nodes; nodes retrorsely pilose; internodes smooth or scabridulous. |
Sheaths | shorter or longer than the internodes, glabrous; ligules 0.8-3 mm, membranous, acute to truncate, erose; blades 0.4-6.5 cm long, 0.5-4.2 mm wide, flat or involute, straight or arcuate-spreading, glabrous abaxially, hirtellous adaxially. |
1.5-8.5 cm, sparsely or densely pilose, hairs to 3 mm, papillose-based; ligules 0.5-0.8 mm, membranous, truncate, erose; blades 2-6 cm long, 1.5-4 mm wide, flat, bases distinctly narrowed to the junction with the sheath, surfaces scabridulous and sparsely pilose, hairs papillose-based. |
Panicles | 1-15 cm long, 0.1-1.7 cm wide, exserted, narrow or spikelike, rachises usually concealed by the branches; primary branches 0.4-5 cm, usually closely appressed at maturity, rarely diverging up to 20° from the rachises; pedicels 0.2-2 mm, setulose. |
6-10.5 cm long, 2.0-4.5 cm wide, secund, open; primary branches 0.8-3.5 cm, secund, spreading at right angles or somewhat reflexed, with 2-5 spikelets; secondary branches not developed; pedicels 1-5 mm, scabrous or shortly pilose, hairs papillose-based; disarticulation at the base of the primary branches. |
Spikelets | 1.7-3.1 mm, occasionally with 2 florets. |
4-8 mm, dimorphic with respect to the glumes, proximal spikelets on each branch almost sessile. |
Glumes | subequal, 0.6-2 mm, 1/3– 1/2 as long as the lemmas, green, 1(2)-veined, acute, sometimes mucronate, mucros less than 0.2 mm; lemmas 1.7-2.6(3.1) mm, lanceolate, dark greenish, plumbeous, or mottled, glabrous, apices scabridulous, acute to acuminate, sometimes mucronate, mucros to 0.5 mm; paleas 1.2-2.4(2.9) mm, lanceolate, acute; anthers 0.9-1.6 mm, yellow to purplish. |
of proximal spikelets on each branch subequal, 0.2-0.7 mm, orbicular, truncate, often erose, unawned; glumes of distal spikelets on each branch markedly unequal; lower glumes to 8 mm, 1-veined, acute, usually awned, awns 0.5-3 mm; upper glumes orbicular, acute, sometimes awn-tipped; lemmas 4.0-7.6 mm, linear to broadly lanceolate, light greenish, smooth or scabrous, usually with greenish veins, apices acuminate, awned, awns 6-19 mm, usually straight, scabrous; paleas 3.7-6.8 mm, narrowly lanceolate, coarsely papillate or almost smooth, acuminate; anthers 0.4-0.8 mm, yellowish. |
Caryopses | 0.9-1.6 mm, narrowly ellipsoid, brown. |
1.8-3 mm, oblong-ovoid, flattened, brownish. |
2n | = 40. |
= 20. |
Muhlenbergia richardsonis |
Muhlenbergia diversiglumis |
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Distribution |
AK; AZ; CA; CO; ID; ME; MI; MN; MT; ND; NE; NM; NV; OR; SD; UT; WA; WI; WY; AB; BC; MB; NB; NT; ON; QC; SK; YT
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Discussion | Muhlenbergia richardsonis grows in open sites in alkaline meadows, prairies, sandy arroyo bottoms, talus slopes, rocky flats and the shores of rivers, at elevations of 60-3300 m. It is the most widespread species of Muhlenbergia in the Flora region, extending from the Yukon Territory to Quebec in the north and to northern Baja California, Mexico, in the south. Morden and Hatch (1996) reported that it also grows in Alaska, but no voucher specimen has been located. Muhlenbergia richardsonis is often confused with M. cuspidata, which differs in lacking rhizomes and having shorter ligules, and sometimes with M. filiformis, which differs in being a weak annual with glabrous internodes and obtuse, erose glumes. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Muhlenbergia diversiglumis has been collected from Galveston County, Texas. The species is native from Mexico to Peru and Venezuela, where it grows on moist cliffs, along water courses, sandy slopes, and road cuts, primarily in moist shaded environments of broadleaf evergreen forests and pine-oak forests, at elevations of 600-2500 m. The collection from Texas may represent a recent introduction. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 25, p. 177. | FNA vol. 25, p. 164. |
Parent taxa | Poaceae > subfam. Chloridoideae > tribe Cynodonteae > Muhlenbergia | Poaceae > subfam. Chloridoideae > tribe Cynodonteae > Muhlenbergia |
Sibling taxa | ||
Synonyms | Sporobolus depauperatus, M. squarrosa | |
Name authority | (Trin.) Rydb. | Trin. |
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