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mat muhly, Matted muhly, muhlenbergie de Richardson, Richardson's muhly, soft-leaf muhly

Navajo muhly, tough muhly

Habit Plants perennial; rhizomatous, not cespitose, often mat-forming. Plants perennial; rhizomatous, rhizomes sometimes short and the plants loosely cespitose.
Culms

5-30 cm tall, 0.4-1 mm thick, decumbent, geniculate, or erect;

internodes usually nodulose (occasionally smooth) for most of their length, puberulent or nodulose below the nodes.

10-50 cm tall, 0.4-1 mm thick, decumbent;

internodes glabrous or strigulose.

Sheaths

shorter or longer than the internodes, glabrous;

ligules 0.8-3 mm, membranous, acute to truncate, erose;

blades 0.4-6.5 cm long, 0.5-4.2 mm wide, flat or involute, straight or arcuate-spreading, glabrous abaxially, hirtellous adaxially.

shorter than the internodes, strigulose or glabrous, usually without necrotic spots, not becoming spirally coiled when old;

ligules 1-2 mm, membranous throughout, obtuse, strigulose or glabrous, erose or toothed, with lateral lobes, lobes less than 1.5 mm longer than the central portion;

blades 1-6 cm long, 1-2 mm wide, flat to involute, smooth or scabridulous abaxially, hirsute adaxially, usually without necrotic spots.

Panicles

1-15 cm long, 0.1-1.7 cm wide, exserted, narrow or spikelike, rachises usually concealed by the branches;

primary branches 0.4-5 cm, usually closely appressed at maturity, rarely diverging up to 20° from the rachises;

pedicels 0.2-2 mm, setulose.

4-13 cm long, 1-3(5) cm wide, not dense;

primary branches 0.5-4 cm, appressed or diverging up to 30° from the rachises, spikelet-bearing to the base;

pedicels 0.1-3 mm.

Spikelets

1.7-3.1 mm, occasionally with 2 florets.

3.5-5 mm.

Glumes

subequal, 0.6-2 mm, 1/3– 1/2 as long as the lemmas, green, 1(2)-veined, acute, sometimes mucronate, mucros less than 0.2 mm;

lemmas 1.7-2.6(3.1) mm, lanceolate, dark greenish, plumbeous, or mottled, glabrous, apices scabridulous, acute to acuminate, sometimes mucronate, mucros to 0.5 mm;

paleas 1.2-2.4(2.9) mm, lanceolate, acute;

anthers 0.9-1.6 mm, yellow to purplish.

subequal, 2-4 mm, exceeded by the florets, 1-veined, scabrous on the veins and near the apices, apices acuminate, unawned or awned, awns to 1.2 mm;

lemmas 3.5-5 mm, lanceolate, mostly purplish, veins conspicuously green, pubescent on the lower 1/3-1/2 of the midveins and margins, hairs to 1.5 mm, apices scabridulous, acuminate, awned, awns 4-12(20) mm, flexuous;

paleas 3.5-5 mm, narrow-lanceolate, intercostal region pubescent, apices acuminate, veins sometimes extending into awns to 0.5 mm;

anthers 1.3-3 mm, purple.

Caryopses

0.9-1.6 mm, narrowly ellipsoid, brown.

2-2.3 mm, fusiform, brownish.

2n

= 40.

= unknown.

Muhlenbergia richardsonis

Muhlenbergia arsenei

Distribution
from FNA
AK; AZ; CA; CO; ID; ME; MI; MN; MT; ND; NE; NM; NV; OR; SD; UT; WA; WI; WY; AB; BC; MB; NB; NT; ON; QC; SK; YT
[WildflowerSearch map]
[BONAP county map]
from FNA
CA; NM; NV; UT
[BONAP county map]
Discussion

Muhlenbergia richardsonis grows in open sites in alkaline meadows, prairies, sandy arroyo bottoms, talus slopes, rocky flats and the shores of rivers, at elevations of 60-3300 m. It is the most widespread species of Muhlenbergia in the Flora region, extending from the Yukon Territory to Quebec in the north and to northern Baja California, Mexico, in the south. Morden and Hatch (1996) reported that it also grows in Alaska, but no voucher specimen has been located.

Muhlenbergia richardsonis is often confused with M. cuspidata, which differs in lacking rhizomes and having shorter ligules, and sometimes with M. filiformis, which differs in being a weak annual with glabrous internodes and obtuse, erose glumes.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Muhlenbergia arsenei grows among granitic boulders, on rocky slopes, limestone rock outcrops, and in arroyos, at elevations of 1400-2850 m. Its range extends from the southwestern United States into Baja California, Mexico. It flowers from August to September.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 25, p. 177. FNA vol. 25, p. 169.
Parent taxa Poaceae > subfam. Chloridoideae > tribe Cynodonteae > Muhlenbergia Poaceae > subfam. Chloridoideae > tribe Cynodonteae > Muhlenbergia
Sibling taxa
M. andina, M. appressa, M. arenacea, M. arenicola, M. arizonica, M. arsenei, M. asperifolia, M. brevis, M. bushii, M. californica, M. capillaris, M. crispiseta, M. curtifolia, M. cuspidata, M. depauperata, M. diversiglumis, M. dubia, M. dumosa, M. elongata, M. eludens, M. emersleyi, M. expansa, M. filiculmis, M. filiformis, M. fragilis, M. frondosa, M. glabrifloris, M. glauca, M. glomerata, M. jonesii, M. lindheimeri, M. longiligula, M. mexicana, M. microsperma, M. minutissima, M. montana, M. palmeri, M. pauciflora, M. pectinata, M. peruviana, M. polycaulis, M. porteri, M. pungens, M. racemosa, M. ramulosa, M. repens, M. reverchonii, M. rigens, M. rigida, M. schreberi, M. sericea, M. setifolia, M. sinuosa, M. sobolifera, M. spiciformis, M. straminea, M. sylvatica, M. tenuiflora, M. tenuifolia, M. texana, M. thurberi, M. torreyana, M. torreyi, M. uniflora, M. utilis, M. villiflora, M. wrightii, M. ×curtisetosa, M. ×involuta
M. andina, M. appressa, M. arenacea, M. arenicola, M. arizonica, M. asperifolia, M. brevis, M. bushii, M. californica, M. capillaris, M. crispiseta, M. curtifolia, M. cuspidata, M. depauperata, M. diversiglumis, M. dubia, M. dumosa, M. elongata, M. eludens, M. emersleyi, M. expansa, M. filiculmis, M. filiformis, M. fragilis, M. frondosa, M. glabrifloris, M. glauca, M. glomerata, M. jonesii, M. lindheimeri, M. longiligula, M. mexicana, M. microsperma, M. minutissima, M. montana, M. palmeri, M. pauciflora, M. pectinata, M. peruviana, M. polycaulis, M. porteri, M. pungens, M. racemosa, M. ramulosa, M. repens, M. reverchonii, M. richardsonis, M. rigens, M. rigida, M. schreberi, M. sericea, M. setifolia, M. sinuosa, M. sobolifera, M. spiciformis, M. straminea, M. sylvatica, M. tenuiflora, M. tenuifolia, M. texana, M. thurberi, M. torreyana, M. torreyi, M. uniflora, M. utilis, M. villiflora, M. wrightii, M. ×curtisetosa, M. ×involuta
Synonyms Sporobolus depauperatus, M. squarrosa
Name authority (Trin.) Rydb. Hitchc.
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