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sandhill muhly

forest muhly, muhlenbergie des bois, woodland muhly

Habit Plants perennial; rhizomatous, not cespitose. Plants perennial; rhizomatous.
Culms

10-70 cm, decumbent below;

internodes cinereous-lanate, glabrous, or scabrous for most of their length, always cinereous-lanate below the nodes.

40-110 cm tall, 1-2 mm thick, erect;

internodes puberulent for most of their length, strigose below the nodes.

Sheaths

longer than the internodes, cinereous-lanate below, glabrous and smooth or scabridulous distally;

ligules 0.2-1 mm, densely ciliate, obtuse, with lateral lobes;

blades 2-8 cm long, 1-2.2 mm wide, flat to tightly involute, scabrous abaxially, hirsute adaxially, stiff, pungent.

glabrous and smooth for most of their length, scabridulous distally, margins hyaline;

ligules 1-2.5 mm, membranous, truncate, lacerate-ciliolate;

blades 5-18 cm long, 3-7 mm wide, flat, scabrous to scabridulous, occasionally smooth.

Panicles

(7)8-16(19) cm long, (2)4-14 cm wide, open;

primary branches 1.5-8 cm, capillary, straight, lower branches diverging 70°-90° from the rachises in mature plants, often appearing fascicled in immature plants;

pedicels 10-25 mm.

terminal and axillary, 6-21 cm long, 0.2-1 cm wide, narrow, not dense;

axillary panicles usually exserted at maturity;

branches 0.8-6 cm, ascending to closely appressed;

pedicels 0.8-3.5 mm, strigose.

Spikelets

2.6-4.5 mm.

2.2-3.7 mm.

Glumes

equal, 1.2-3 mm, purplish near the base, smooth or scabridulous distally, 1-veined, acuminate or acute, unawned or awned, awns to 1 mm;

lemmas 2.6-4.5 mm, lanceolate, purplish, scabridulous distally and on the margins, apices acuminate, awned, awns 1-1.5(2) mm, straight;

paleas 2.6-4.5 mm, lanceolate, glabrous, acuminate, 2-awned, awns to 1 mm;

anthers 1.8-2.6 mm, purplish.

subequal, 1.8-3 mm, nearly as long as the lemmas, 1-veined, tapering from near the base, apices scabridulous, acuminate, unawned or awned, awns to 1 mm;

lemmas 2.2-3.7 mm, lanceolate to narrowly lanceolate, hairy on the calluses, lower 1/2 of the midveins, and margins, hairs 0.2-0.5 mm, apices scabridulous, acuminate, awned, awns 5-18 mm, purplish;

paleas 2-3.5 mm, lanceolate, proximal 1/2 shortly pilose, apices scabridulous, acuminate;

anthers 0.4-0.8 mm, yellow.

Caryopses

1.8-2.5 mm, fusiform, brownish.

1.4-2 mm, fusiform, brown.

2n

= 26, 42, 60.

= 40.

Muhlenbergia pungens

Muhlenbergia sylvatica

Distribution
from FNA
AZ; CO; NE; NM; SD; TX; UT; WY
[WildflowerSearch map]
[BONAP county map]
from FNA
AL; AR; AZ; CT; DC; DE; GA; IA; IL; IN; KS; KY; MA; MD; ME; MI; MN; MO; MS; NC; NE; NH; NJ; NY; OH; OK; PA; RI; SC; SD; TN; TX; VA; VT; WI; WV; ON; QC
[WildflowerSearch map]
[BONAP county map]
Discussion

Muhlenbergia pungens grows in loose sandy soils near sand dunes to sandy clay loam slopes and flats in desert shrub and open woodlands, at elevations of 600-2500 m. It is known only from the western and central contiguous United States.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Muhlenbergia sylvatica grows in upland forests, along creeks and hollows, on rocky ledges derived from sandstone, shale, or calcareous parent materials, moist prairies, and swamps, at elevations from 30-1500 m. It is restricted to the Flora region, its primary range being southeastern Canada and the midwestern and eastern United States. Reports from British Columbia were based on a misidentification (Douglas et al. 2002). The record from Arizona is based on the report in Kearney and Peebles (1951) of a collection made by Toumey at Grapevine Creek in the Grand Canyon.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 25, p. 173. FNA vol. 25, p. 160.
Parent taxa Poaceae > subfam. Chloridoideae > tribe Cynodonteae > Muhlenbergia Poaceae > subfam. Chloridoideae > tribe Cynodonteae > Muhlenbergia
Sibling taxa
M. andina, M. appressa, M. arenacea, M. arenicola, M. arizonica, M. arsenei, M. asperifolia, M. brevis, M. bushii, M. californica, M. capillaris, M. crispiseta, M. curtifolia, M. cuspidata, M. depauperata, M. diversiglumis, M. dubia, M. dumosa, M. elongata, M. eludens, M. emersleyi, M. expansa, M. filiculmis, M. filiformis, M. fragilis, M. frondosa, M. glabrifloris, M. glauca, M. glomerata, M. jonesii, M. lindheimeri, M. longiligula, M. mexicana, M. microsperma, M. minutissima, M. montana, M. palmeri, M. pauciflora, M. pectinata, M. peruviana, M. polycaulis, M. porteri, M. racemosa, M. ramulosa, M. repens, M. reverchonii, M. richardsonis, M. rigens, M. rigida, M. schreberi, M. sericea, M. setifolia, M. sinuosa, M. sobolifera, M. spiciformis, M. straminea, M. sylvatica, M. tenuiflora, M. tenuifolia, M. texana, M. thurberi, M. torreyana, M. torreyi, M. uniflora, M. utilis, M. villiflora, M. wrightii, M. ×curtisetosa, M. ×involuta
M. andina, M. appressa, M. arenacea, M. arenicola, M. arizonica, M. arsenei, M. asperifolia, M. brevis, M. bushii, M. californica, M. capillaris, M. crispiseta, M. curtifolia, M. cuspidata, M. depauperata, M. diversiglumis, M. dubia, M. dumosa, M. elongata, M. eludens, M. emersleyi, M. expansa, M. filiculmis, M. filiformis, M. fragilis, M. frondosa, M. glabrifloris, M. glauca, M. glomerata, M. jonesii, M. lindheimeri, M. longiligula, M. mexicana, M. microsperma, M. minutissima, M. montana, M. palmeri, M. pauciflora, M. pectinata, M. peruviana, M. polycaulis, M. porteri, M. pungens, M. racemosa, M. ramulosa, M. repens, M. reverchonii, M. richardsonis, M. rigens, M. rigida, M. schreberi, M. sericea, M. setifolia, M. sinuosa, M. sobolifera, M. spiciformis, M. straminea, M. tenuiflora, M. tenuifolia, M. texana, M. thurberi, M. torreyana, M. torreyi, M. uniflora, M. utilis, M. villiflora, M. wrightii, M. ×curtisetosa, M. ×involuta
Synonyms M. sylvatica var. robusta, M. sylvatica forma attenuata
Name authority Thurb. ex A. Gray (Torn) Torr. ex A. Gray
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