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alkali muhly, or alkali muhly, scratch grass

mixedglume muhly

Habit Plants perennial; rhizomatous, not cespitose, occasionally: stoloniferous. Plants annual; sprawling.
Culms

10-60(100) cm, decumbent-ascending, bases somewhat compressed-keeled;

internodes glabrous, shiny below the nodes.

16-50 cm, decumbent, rooting at the lower nodes;

nodes retrorsely pilose;

internodes smooth or scabridulous.

Sheaths

glabrous, margins hyaline;

ligules 0.2-1 mm, firm, truncate, ciliate, without lateral lobes;

blades 2-7(11) cm long, 1-2.8(4) mm wide, flat, occasionally conduplicate, smooth or scabridulous abaxially, scabridulous adaxially, margins and midveins not conspicuously thickened, greenish, apices acute, not sharp.

1.5-8.5 cm, sparsely or densely pilose, hairs to 3 mm, papillose-based;

ligules 0.5-0.8 mm, membranous, truncate, erose;

blades 2-6 cm long, 1.5-4 mm wide, flat, bases distinctly narrowed to the junction with the sheath, surfaces scabridulous and sparsely pilose, hairs papillose-based.

Panicles

6-21 cm long, 4-16 cm wide, broadly ovoid, open;

primary branches 3-12 cm, capillary, lower branches spreading 30-90° from the rachises, never appearing fascicled;

pedicels 3-14 mm, longer than the spikelets.

6-10.5 cm long, 2.0-4.5 cm wide, secund, open;

primary branches 0.8-3.5 cm, secund, spreading at right angles or somewhat reflexed, with 2-5 spikelets;

secondary branches not developed;

pedicels 1-5 mm, scabrous or shortly pilose, hairs papillose-based;

disarticulation at the base of the primary branches.

Spikelets

1.2-2.1 mm, occasionally with 2 or 3 florets.

4-8 mm, dimorphic with respect to the glumes, proximal spikelets on each branch almost sessile.

Glumes

equal, 0.6-1.7 mm, purplish, scabridulous, particularly on the veins, 1-veined, apices acute;

lemmas 1.2-2.1 mm, lanceolate to oblong-elliptic, somewhat plumbeous, glabrous, usually smooth, occasionally scabridulous near the apices, apices acute, unawned or mucronate, mucros to 0.3 mm;

paleas 1.2-2.1 mm, lanceolate, glabrous, acute;

anthers 1-1.3 mm, greenish-yellow to purplish at maturity.

of proximal spikelets on each branch subequal, 0.2-0.7 mm, orbicular, truncate, often erose, unawned;

glumes of distal spikelets on each branch markedly unequal;

lower glumes to 8 mm, 1-veined, acute, usually awned, awns 0.5-3 mm;

upper glumes orbicular, acute, sometimes awn-tipped;

lemmas 4.0-7.6 mm, linear to broadly lanceolate, light greenish, smooth or scabrous, usually with greenish veins, apices acuminate, awned, awns 6-19 mm, usually straight, scabrous;

paleas 3.7-6.8 mm, narrowly lanceolate, coarsely papillate or almost smooth, acuminate;

anthers 0.4-0.8 mm, yellowish.

Caryopses

0.8-1 mm, fusiform, brownish.

1.8-3 mm, oblong-ovoid, flattened, brownish.

2n

= 20, 22, 28.

= 20.

Muhlenbergia asperifolia

Muhlenbergia diversiglumis

Distribution
from FNA
AZ; CA; CO; IA; ID; IL; IN; KS; KY; MD; MI; MN; MO; MT; ND; NE; NM; NV; NY; OH; OK; OR; PA; SD; TX; UT; WA; WI; WY; AB; BC; MB; ON; SK
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Discussion

Muhlenbergia asperifolia grows in moist, often alkaline meadows, playa margins, and sandy washes, on grassy slopes, and around seeps and hot springs, at elevations of 55-3000 m. Its geographic range includes northern Mexico. Muhlenbergia asperifolia is morphologically similar to the southeastern M. torreyana, but differs in having glabrous, weakly compressed culms and more widely divergent panicle branches.

The caryopses of Muhlenbergia asperifolia are frequently infected by a smut, Tilletia asperifolia Ellis & Everhart, which produces a globose body filled with blackish-brown spores.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Muhlenbergia diversiglumis has been collected from Galveston County, Texas. The species is native from Mexico to Peru and Venezuela, where it grows on moist cliffs, along water courses, sandy slopes, and road cuts, primarily in moist shaded environments of broadleaf evergreen forests and pine-oak forests, at elevations of 600-2500 m. The collection from Texas may represent a recent introduction.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 25, p. 179. FNA vol. 25, p. 164.
Parent taxa Poaceae > subfam. Chloridoideae > tribe Cynodonteae > Muhlenbergia Poaceae > subfam. Chloridoideae > tribe Cynodonteae > Muhlenbergia
Sibling taxa
M. andina, M. appressa, M. arenacea, M. arenicola, M. arizonica, M. arsenei, M. brevis, M. bushii, M. californica, M. capillaris, M. crispiseta, M. curtifolia, M. cuspidata, M. depauperata, M. diversiglumis, M. dubia, M. dumosa, M. elongata, M. eludens, M. emersleyi, M. expansa, M. filiculmis, M. filiformis, M. fragilis, M. frondosa, M. glabrifloris, M. glauca, M. glomerata, M. jonesii, M. lindheimeri, M. longiligula, M. mexicana, M. microsperma, M. minutissima, M. montana, M. palmeri, M. pauciflora, M. pectinata, M. peruviana, M. polycaulis, M. porteri, M. pungens, M. racemosa, M. ramulosa, M. repens, M. reverchonii, M. richardsonis, M. rigens, M. rigida, M. schreberi, M. sericea, M. setifolia, M. sinuosa, M. sobolifera, M. spiciformis, M. straminea, M. sylvatica, M. tenuiflora, M. tenuifolia, M. texana, M. thurberi, M. torreyana, M. torreyi, M. uniflora, M. utilis, M. villiflora, M. wrightii, M. ×curtisetosa, M. ×involuta
M. andina, M. appressa, M. arenacea, M. arenicola, M. arizonica, M. arsenei, M. asperifolia, M. brevis, M. bushii, M. californica, M. capillaris, M. crispiseta, M. curtifolia, M. cuspidata, M. depauperata, M. dubia, M. dumosa, M. elongata, M. eludens, M. emersleyi, M. expansa, M. filiculmis, M. filiformis, M. fragilis, M. frondosa, M. glabrifloris, M. glauca, M. glomerata, M. jonesii, M. lindheimeri, M. longiligula, M. mexicana, M. microsperma, M. minutissima, M. montana, M. palmeri, M. pauciflora, M. pectinata, M. peruviana, M. polycaulis, M. porteri, M. pungens, M. racemosa, M. ramulosa, M. repens, M. reverchonii, M. richardsonis, M. rigens, M. rigida, M. schreberi, M. sericea, M. setifolia, M. sinuosa, M. sobolifera, M. spiciformis, M. straminea, M. sylvatica, M. tenuiflora, M. tenuifolia, M. texana, M. thurberi, M. torreyana, M. torreyi, M. uniflora, M. utilis, M. villiflora, M. wrightii, M. ×curtisetosa, M. ×involuta
Name authority (Nees & Meyen ex Trin.) Parodi Trin.
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