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alkali muhly, or alkali muhly, scratch grass

California muhly

Habit Plants perennial; rhizomatous, not cespitose, occasionally: stoloniferous. Plants perennial; rhizomatous, not cespitose.
Culms

10-60(100) cm, decumbent-ascending, bases somewhat compressed-keeled;

internodes glabrous, shiny below the nodes.

30-70 cm tall, 0.7-1.5 mm thick, decumbent;

internodes dull, smooth, and glabrous for most of their length, sometimes strigose immediately below the nodes.

Sheaths

glabrous, margins hyaline;

ligules 0.2-1 mm, firm, truncate, ciliate, without lateral lobes;

blades 2-7(11) cm long, 1-2.8(4) mm wide, flat, occasionally conduplicate, smooth or scabridulous abaxially, scabridulous adaxially, margins and midveins not conspicuously thickened, greenish, apices acute, not sharp.

shorter than the internodes, scabrous, margins whitish;

ligules 0.8-2 mm, membranous, truncate, ciliolate, irregularly toothed;

blades 4-16 cm long, 2-6 mm wide, flat, scabridulous abaxially, scabrous to strigose adaxially.

Panicles

6-21 cm long, 4-16 cm wide, broadly ovoid, open;

primary branches 3-12 cm, capillary, lower branches spreading 30-90° from the rachises, never appearing fascicled;

pedicels 3-14 mm, longer than the spikelets.

terminal, 5-13 cm long, 0.5-2.2 cm wide, dense;

branches 0.5-3.2 cm, ascending, appressed or diverging up to 20° from the rachises;

pedicels to 1.5 mm, stout, strigose;

axillary panicles not present.

Spikelets

1.2-2.1 mm, occasionally with 2 or 3 florets.

2.8-4 mm.

Glumes

equal, 0.6-1.7 mm, purplish, scabridulous, particularly on the veins, 1-veined, apices acute;

lemmas 1.2-2.1 mm, lanceolate to oblong-elliptic, somewhat plumbeous, glabrous, usually smooth, occasionally scabridulous near the apices, apices acute, unawned or mucronate, mucros to 0.3 mm;

paleas 1.2-2.1 mm, lanceolate, glabrous, acute;

anthers 1-1.3 mm, greenish-yellow to purplish at maturity.

subequal, 2.5-4 mm, nearly as long as or slightly longer than the lemmas, scabrous (especially on the veins), 1-veined, tapering from the base to the acuminate apices, usually unawned, awns, if present, to 1.2 mm;

lemmas 2.8-4 mm, narrowly lanceolate, with soft hairs on the calluses and lower portion of the lemma bodies, hairs to 1 mm, apices scabridulous, acuminate, awned, awns 0.2-2.2 mm;

paleas 2.8-4 mm, subequal to the lemmas, narrowly lanceolate, with short (less than 1.5 mm), soft hairs on the lower 1/2, apices scabridulous, acuminate;

anthers 1-1.7 mm, yellow.

Caryopses

0.8-1 mm, fusiform, brownish.

1.7-2 mm, fusiform, brown.

2n

= 20, 22, 28.

= 80.

Muhlenbergia asperifolia

Muhlenbergia californica

Distribution
from FNA
AZ; CA; CO; IA; ID; IL; IN; KS; KY; MD; MI; MN; MO; MT; ND; NE; NM; NV; NY; OH; OK; OR; PA; SD; TX; UT; WA; WI; WY; AB; BC; MB; ON; SK
[WildflowerSearch map]
[BONAP county map]
from FNA
CA
[BONAP county map]
Discussion

Muhlenbergia asperifolia grows in moist, often alkaline meadows, playa margins, and sandy washes, on grassy slopes, and around seeps and hot springs, at elevations of 55-3000 m. Its geographic range includes northern Mexico. Muhlenbergia asperifolia is morphologically similar to the southeastern M. torreyana, but differs in having glabrous, weakly compressed culms and more widely divergent panicle branches.

The caryopses of Muhlenbergia asperifolia are frequently infected by a smut, Tilletia asperifolia Ellis & Everhart, which produces a globose body filled with blackish-brown spores.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Muhlenbergia californica grows in canyons, along moist ditches, and on sandy slopes, at elevations of 100-2150 m. It is endemic to the Transverse Ranges of southern California.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 25, p. 179. FNA vol. 25, p. 154.
Parent taxa Poaceae > subfam. Chloridoideae > tribe Cynodonteae > Muhlenbergia Poaceae > subfam. Chloridoideae > tribe Cynodonteae > Muhlenbergia
Sibling taxa
M. andina, M. appressa, M. arenacea, M. arenicola, M. arizonica, M. arsenei, M. brevis, M. bushii, M. californica, M. capillaris, M. crispiseta, M. curtifolia, M. cuspidata, M. depauperata, M. diversiglumis, M. dubia, M. dumosa, M. elongata, M. eludens, M. emersleyi, M. expansa, M. filiculmis, M. filiformis, M. fragilis, M. frondosa, M. glabrifloris, M. glauca, M. glomerata, M. jonesii, M. lindheimeri, M. longiligula, M. mexicana, M. microsperma, M. minutissima, M. montana, M. palmeri, M. pauciflora, M. pectinata, M. peruviana, M. polycaulis, M. porteri, M. pungens, M. racemosa, M. ramulosa, M. repens, M. reverchonii, M. richardsonis, M. rigens, M. rigida, M. schreberi, M. sericea, M. setifolia, M. sinuosa, M. sobolifera, M. spiciformis, M. straminea, M. sylvatica, M. tenuiflora, M. tenuifolia, M. texana, M. thurberi, M. torreyana, M. torreyi, M. uniflora, M. utilis, M. villiflora, M. wrightii, M. ×curtisetosa, M. ×involuta
M. andina, M. appressa, M. arenacea, M. arenicola, M. arizonica, M. arsenei, M. asperifolia, M. brevis, M. bushii, M. capillaris, M. crispiseta, M. curtifolia, M. cuspidata, M. depauperata, M. diversiglumis, M. dubia, M. dumosa, M. elongata, M. eludens, M. emersleyi, M. expansa, M. filiculmis, M. filiformis, M. fragilis, M. frondosa, M. glabrifloris, M. glauca, M. glomerata, M. jonesii, M. lindheimeri, M. longiligula, M. mexicana, M. microsperma, M. minutissima, M. montana, M. palmeri, M. pauciflora, M. pectinata, M. peruviana, M. polycaulis, M. porteri, M. pungens, M. racemosa, M. ramulosa, M. repens, M. reverchonii, M. richardsonis, M. rigens, M. rigida, M. schreberi, M. sericea, M. setifolia, M. sinuosa, M. sobolifera, M. spiciformis, M. straminea, M. sylvatica, M. tenuiflora, M. tenuifolia, M. texana, M. thurberi, M. torreyana, M. torreyi, M. uniflora, M. utilis, M. villiflora, M. wrightii, M. ×curtisetosa, M. ×involuta
Name authority (Nees & Meyen ex Trin.) Parodi Vasey
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