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alkali muhly, or alkali muhly, scratch grass

Navajo muhly, tough muhly

Habit Plants perennial; rhizomatous, not cespitose, occasionally: stoloniferous. Plants perennial; rhizomatous, rhizomes sometimes short and the plants loosely cespitose.
Culms

10-60(100) cm, decumbent-ascending, bases somewhat compressed-keeled;

internodes glabrous, shiny below the nodes.

10-50 cm tall, 0.4-1 mm thick, decumbent;

internodes glabrous or strigulose.

Sheaths

glabrous, margins hyaline;

ligules 0.2-1 mm, firm, truncate, ciliate, without lateral lobes;

blades 2-7(11) cm long, 1-2.8(4) mm wide, flat, occasionally conduplicate, smooth or scabridulous abaxially, scabridulous adaxially, margins and midveins not conspicuously thickened, greenish, apices acute, not sharp.

shorter than the internodes, strigulose or glabrous, usually without necrotic spots, not becoming spirally coiled when old;

ligules 1-2 mm, membranous throughout, obtuse, strigulose or glabrous, erose or toothed, with lateral lobes, lobes less than 1.5 mm longer than the central portion;

blades 1-6 cm long, 1-2 mm wide, flat to involute, smooth or scabridulous abaxially, hirsute adaxially, usually without necrotic spots.

Panicles

6-21 cm long, 4-16 cm wide, broadly ovoid, open;

primary branches 3-12 cm, capillary, lower branches spreading 30-90° from the rachises, never appearing fascicled;

pedicels 3-14 mm, longer than the spikelets.

4-13 cm long, 1-3(5) cm wide, not dense;

primary branches 0.5-4 cm, appressed or diverging up to 30° from the rachises, spikelet-bearing to the base;

pedicels 0.1-3 mm.

Spikelets

1.2-2.1 mm, occasionally with 2 or 3 florets.

3.5-5 mm.

Glumes

equal, 0.6-1.7 mm, purplish, scabridulous, particularly on the veins, 1-veined, apices acute;

lemmas 1.2-2.1 mm, lanceolate to oblong-elliptic, somewhat plumbeous, glabrous, usually smooth, occasionally scabridulous near the apices, apices acute, unawned or mucronate, mucros to 0.3 mm;

paleas 1.2-2.1 mm, lanceolate, glabrous, acute;

anthers 1-1.3 mm, greenish-yellow to purplish at maturity.

subequal, 2-4 mm, exceeded by the florets, 1-veined, scabrous on the veins and near the apices, apices acuminate, unawned or awned, awns to 1.2 mm;

lemmas 3.5-5 mm, lanceolate, mostly purplish, veins conspicuously green, pubescent on the lower 1/3-1/2 of the midveins and margins, hairs to 1.5 mm, apices scabridulous, acuminate, awned, awns 4-12(20) mm, flexuous;

paleas 3.5-5 mm, narrow-lanceolate, intercostal region pubescent, apices acuminate, veins sometimes extending into awns to 0.5 mm;

anthers 1.3-3 mm, purple.

Caryopses

0.8-1 mm, fusiform, brownish.

2-2.3 mm, fusiform, brownish.

2n

= 20, 22, 28.

= unknown.

Muhlenbergia asperifolia

Muhlenbergia arsenei

Distribution
from FNA
AZ; CA; CO; IA; ID; IL; IN; KS; KY; MD; MI; MN; MO; MT; ND; NE; NM; NV; NY; OH; OK; OR; PA; SD; TX; UT; WA; WI; WY; AB; BC; MB; ON; SK
[WildflowerSearch map]
[BONAP county map]
from FNA
CA; NM; NV; UT
[BONAP county map]
Discussion

Muhlenbergia asperifolia grows in moist, often alkaline meadows, playa margins, and sandy washes, on grassy slopes, and around seeps and hot springs, at elevations of 55-3000 m. Its geographic range includes northern Mexico. Muhlenbergia asperifolia is morphologically similar to the southeastern M. torreyana, but differs in having glabrous, weakly compressed culms and more widely divergent panicle branches.

The caryopses of Muhlenbergia asperifolia are frequently infected by a smut, Tilletia asperifolia Ellis & Everhart, which produces a globose body filled with blackish-brown spores.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Muhlenbergia arsenei grows among granitic boulders, on rocky slopes, limestone rock outcrops, and in arroyos, at elevations of 1400-2850 m. Its range extends from the southwestern United States into Baja California, Mexico. It flowers from August to September.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 25, p. 179. FNA vol. 25, p. 169.
Parent taxa Poaceae > subfam. Chloridoideae > tribe Cynodonteae > Muhlenbergia Poaceae > subfam. Chloridoideae > tribe Cynodonteae > Muhlenbergia
Sibling taxa
M. andina, M. appressa, M. arenacea, M. arenicola, M. arizonica, M. arsenei, M. brevis, M. bushii, M. californica, M. capillaris, M. crispiseta, M. curtifolia, M. cuspidata, M. depauperata, M. diversiglumis, M. dubia, M. dumosa, M. elongata, M. eludens, M. emersleyi, M. expansa, M. filiculmis, M. filiformis, M. fragilis, M. frondosa, M. glabrifloris, M. glauca, M. glomerata, M. jonesii, M. lindheimeri, M. longiligula, M. mexicana, M. microsperma, M. minutissima, M. montana, M. palmeri, M. pauciflora, M. pectinata, M. peruviana, M. polycaulis, M. porteri, M. pungens, M. racemosa, M. ramulosa, M. repens, M. reverchonii, M. richardsonis, M. rigens, M. rigida, M. schreberi, M. sericea, M. setifolia, M. sinuosa, M. sobolifera, M. spiciformis, M. straminea, M. sylvatica, M. tenuiflora, M. tenuifolia, M. texana, M. thurberi, M. torreyana, M. torreyi, M. uniflora, M. utilis, M. villiflora, M. wrightii, M. ×curtisetosa, M. ×involuta
M. andina, M. appressa, M. arenacea, M. arenicola, M. arizonica, M. asperifolia, M. brevis, M. bushii, M. californica, M. capillaris, M. crispiseta, M. curtifolia, M. cuspidata, M. depauperata, M. diversiglumis, M. dubia, M. dumosa, M. elongata, M. eludens, M. emersleyi, M. expansa, M. filiculmis, M. filiformis, M. fragilis, M. frondosa, M. glabrifloris, M. glauca, M. glomerata, M. jonesii, M. lindheimeri, M. longiligula, M. mexicana, M. microsperma, M. minutissima, M. montana, M. palmeri, M. pauciflora, M. pectinata, M. peruviana, M. polycaulis, M. porteri, M. pungens, M. racemosa, M. ramulosa, M. repens, M. reverchonii, M. richardsonis, M. rigens, M. rigida, M. schreberi, M. sericea, M. setifolia, M. sinuosa, M. sobolifera, M. spiciformis, M. straminea, M. sylvatica, M. tenuiflora, M. tenuifolia, M. texana, M. thurberi, M. torreyana, M. torreyi, M. uniflora, M. utilis, M. villiflora, M. wrightii, M. ×curtisetosa, M. ×involuta
Name authority (Nees & Meyen ex Trin.) Parodi Hitchc.
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