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alkali muhly, or alkali muhly, scratch grass

appressed muhly, devil's-canyon muhly, spreading littleseed muhly

Habit Plants perennial; rhizomatous, not cespitose, occasionally: stoloniferous. Plants annual.
Culms

10-60(100) cm, decumbent-ascending, bases somewhat compressed-keeled;

internodes glabrous, shiny below the nodes.

10-42 cm, erect or decumbent, much branched below;

internodes mostly scabrous, scabrous or hispidulous below the nodes.

Sheaths

glabrous, margins hyaline;

ligules 0.2-1 mm, firm, truncate, ciliate, without lateral lobes;

blades 2-7(11) cm long, 1-2.8(4) mm wide, flat, occasionally conduplicate, smooth or scabridulous abaxially, scabridulous adaxially, margins and midveins not conspicuously thickened, greenish, apices acute, not sharp.

usually shorter than the internodes, flattened below, glabrous, scabridulous or smooth, striate;

ligules 1.5-3 mm, hyaline, acute, lacerate, with lateral lobes;

blades 1-5(7) cm long, 0.8-1.6(2) mm wide, flat or involute, scabrous abaxially, pubescent adaxially.

Panicles

6-21 cm long, 4-16 cm wide, broadly ovoid, open;

primary branches 3-12 cm, capillary, lower branches spreading 30-90° from the rachises, never appearing fascicled;

pedicels 3-14 mm, longer than the spikelets.

4-23 cm long, 0.5-2 cm wide, usually partially included in the subtending sheaths at maturity;

primary branches 20-45 mm, ascending, closely appressed;

pedicels 1-5 mm, appressed, scabrous;

disarticulation above the glumes.

Spikelets

1.2-2.1 mm, occasionally with 2 or 3 florets.

4-7.5 mm, borne singly.

Glumes

equal, 0.6-1.7 mm, purplish, scabridulous, particularly on the veins, 1-veined, apices acute;

lemmas 1.2-2.1 mm, lanceolate to oblong-elliptic, somewhat plumbeous, glabrous, usually smooth, occasionally scabridulous near the apices, apices acute, unawned or mucronate, mucros to 0.3 mm;

paleas 1.2-2.1 mm, lanceolate, glabrous, acute;

anthers 1-1.3 mm, greenish-yellow to purplish at maturity.

1-2 mm, 1-veined, veins conspicuous, scabrous, and greenish basally, apices obtuse to subacute, often erose;

lemmas 4-6.2(7.5) mm, narrowly lanceolate, terete, mostly smooth, scabrous distally, hairy on the calluses and lower 1/4 of the margins, hairs 0.2-0.3 mm, whitish, lemma bodies not appearing 5-veined, apices acuminate, awned, awns 10-30 mm, straight, scabrous;

paleas 3.8-5.7(7) mm, narrowly lanceolate;

anthers 0.3-1.1 mm, purplish.

Caryopses

0.8-1 mm, fusiform, brownish.

1.6-3 mm, narrowly fusiform, brownish to pinkish.

Cleistogamous

spikelets usually present in the axils of the lower leaves.

2n

= 20, 22, 28.

= unknown.

Muhlenbergia asperifolia

Muhlenbergia appressa

Distribution
from FNA
AZ; CA; CO; IA; ID; IL; IN; KS; KY; MD; MI; MN; MO; MT; ND; NE; NM; NV; NY; OH; OK; OR; PA; SD; TX; UT; WA; WI; WY; AB; BC; MB; ON; SK
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; CA
[WildflowerSearch map]
[BONAP county map]
Discussion

Muhlenbergia asperifolia grows in moist, often alkaline meadows, playa margins, and sandy washes, on grassy slopes, and around seeps and hot springs, at elevations of 55-3000 m. Its geographic range includes northern Mexico. Muhlenbergia asperifolia is morphologically similar to the southeastern M. torreyana, but differs in having glabrous, weakly compressed culms and more widely divergent panicle branches.

The caryopses of Muhlenbergia asperifolia are frequently infected by a smut, Tilletia asperifolia Ellis & Everhart, which produces a globose body filled with blackish-brown spores.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Muhlenbergia appressa grows in sandy drainages, canyon bottoms, rocky road cuts, and sandy slopes, at elevations of 20-1750 m. Its range extends from Arizona to Baja California, Mexico. It grows in gramma grasslands, oak-juniper woodlands, and chaparral associations.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 25, p. 179. FNA vol. 25, p. 164.
Parent taxa Poaceae > subfam. Chloridoideae > tribe Cynodonteae > Muhlenbergia Poaceae > subfam. Chloridoideae > tribe Cynodonteae > Muhlenbergia
Sibling taxa
M. andina, M. appressa, M. arenacea, M. arenicola, M. arizonica, M. arsenei, M. brevis, M. bushii, M. californica, M. capillaris, M. crispiseta, M. curtifolia, M. cuspidata, M. depauperata, M. diversiglumis, M. dubia, M. dumosa, M. elongata, M. eludens, M. emersleyi, M. expansa, M. filiculmis, M. filiformis, M. fragilis, M. frondosa, M. glabrifloris, M. glauca, M. glomerata, M. jonesii, M. lindheimeri, M. longiligula, M. mexicana, M. microsperma, M. minutissima, M. montana, M. palmeri, M. pauciflora, M. pectinata, M. peruviana, M. polycaulis, M. porteri, M. pungens, M. racemosa, M. ramulosa, M. repens, M. reverchonii, M. richardsonis, M. rigens, M. rigida, M. schreberi, M. sericea, M. setifolia, M. sinuosa, M. sobolifera, M. spiciformis, M. straminea, M. sylvatica, M. tenuiflora, M. tenuifolia, M. texana, M. thurberi, M. torreyana, M. torreyi, M. uniflora, M. utilis, M. villiflora, M. wrightii, M. ×curtisetosa, M. ×involuta
M. andina, M. arenacea, M. arenicola, M. arizonica, M. arsenei, M. asperifolia, M. brevis, M. bushii, M. californica, M. capillaris, M. crispiseta, M. curtifolia, M. cuspidata, M. depauperata, M. diversiglumis, M. dubia, M. dumosa, M. elongata, M. eludens, M. emersleyi, M. expansa, M. filiculmis, M. filiformis, M. fragilis, M. frondosa, M. glabrifloris, M. glauca, M. glomerata, M. jonesii, M. lindheimeri, M. longiligula, M. mexicana, M. microsperma, M. minutissima, M. montana, M. palmeri, M. pauciflora, M. pectinata, M. peruviana, M. polycaulis, M. porteri, M. pungens, M. racemosa, M. ramulosa, M. repens, M. reverchonii, M. richardsonis, M. rigens, M. rigida, M. schreberi, M. sericea, M. setifolia, M. sinuosa, M. sobolifera, M. spiciformis, M. straminea, M. sylvatica, M. tenuiflora, M. tenuifolia, M. texana, M. thurberi, M. torreyana, M. torreyi, M. uniflora, M. utilis, M. villiflora, M. wrightii, M. ×curtisetosa, M. ×involuta
Name authority (Nees & Meyen ex Trin.) Parodi CO. Goodd.
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