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maidenhair vine, wirevine

wirevine

Habit Shrubs, vinelike, perennial; rhizomatous.
Stems

suberect, prostrate, or scandent, glabrous or puberulent, sometimes papillose.

Leaves

deciduous, cauline, alternate, usually petiolate;

ocrea usually deciduous, sometimes persistent, chartaceous;

petiole base articulated, extrafloral nectaries present;

blade linear to orbiculate, panduriform, or triangular-lanceolate, margins entire or irregularly wavy.

Inflorescences

terminal and axillary, spikelike, essentially not pedunculate.

Pedicels

prescent.

Flowers

bisexual and unisexual, with staminate, pistillate, or sometimes both sexes occurring with bisexual flowers on the same plant, 1–2(–5) per ocreate fascicle, base stipelike;

perianth accrescent, white to greenish white, campanulate, glabrous;

tepals 5, connate proximally, sepaloid, dimorphic, outer slightly larger than inner.

Staminate flowers

stamens 8 (9);

filaments distinct, adnate to base of perianth tube, glabrous;

anthers yellow or pink to purple, ovate to elliptic;

pistil rudimentary.

Pistillate flowers

tube white or reddish purple to black in fruit, becoming fleshy;

stamens rudimentary;

styles 3, spreading, connate proximally;

stigmas fimbriate.

Achenes

completely or partly included in fleshy perianth, black or dark brown, unwinged, 3-gonous to subglobose, glabrous.

Seeds

embryo straight.

x

= 10.

Muehlenbeckia

Muehlenbeckia hastulata

Distribution
from USDA
Central America; South America; Pacific Islands (New Zealand); Australia [Introduced in North America]
[BONAP county map]
from FNA
CA; South America
[BONAP county map]
Discussion

species 23 (2 in the flora)

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 3 (1 in the flora).

J. Brandbyge (1992) recognized two other varieties of Muehlenbeckia hastulata distinguished by leaf shape, inflorescence size, and achene morphology. Variety rotundata (Philippi) Brandbyge is endemic in Argentina and Chile; var. fascicularis (Meisner) Brandbyge is endemic in Chile.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Tubes of pistillate flowers white in fruit; distal branches of stems brownish-puberulent; leaf blades ovate-oblong to suborbiculate or panduriform
M. complexa
1. Tubes of pistillate flowers reddish purple to black in fruit; distal branches of stems glabrous; leaf blades triangular-lanceolate
M. hastulata
Source FNA vol. 5, p. 485. Author: Craig C. Freeman. FNA vol. 5, p. 486.
Parent taxa Polygonaceae > subfam. Polygonoideae Polygonaceae > subfam. Polygonoideae > Muehlenbeckia
Sibling taxa
M. complexa
Subordinate taxa
M. complexa, M. hastulata
M. hastulata var. hastulata
Synonyms Rumex hastulatus
Name authority Meisner: Pl. Vasc. Gen. 1: 316 (Smith) I. M. Johnston: Contr. Gray Herb. 81: 88. (1928)
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