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maidenhair vine, wirevine

lacy wirevine, maidenhair vine, Mattress vine

Habit Shrubs, vinelike, perennial; rhizomatous. Plants (0.5–)1–5 m. Stems prostrate to scandent, especially distally, sometimes rooting at nodes, striate, branched, glabrescent or puberulent, distal branches brownish-puberulent.
Stems

suberect, prostrate, or scandent, glabrous or puberulent, sometimes papillose.

Leaves

deciduous, cauline, alternate, usually petiolate;

ocrea usually deciduous, sometimes persistent, chartaceous;

petiole base articulated, extrafloral nectaries present;

blade linear to orbiculate, panduriform, or triangular-lanceolate, margins entire or irregularly wavy.

ocrea deciduous, brownish hyaline, cylindric, 2–3 mm, margins truncate, eciliate, faces puberulent along veins;

petiole 3–10 mm, brownish-puberulent;

blade ovate-oblong to suborbiculate or panduriform, 0.5–2.5(–4) × 0.5–2.5(–4) cm, coriaceous, base truncate, margins entire, glabrous or scabrous, apex rounded to apiculate, glabrous adaxially and abaxially, or puberulent abaxially along midvein, obscurely punctuate abaxially.

Inflorescences

terminal and axillary, spikelike, essentially not pedunculate.

terminal and axillary, 5–30 mm.

Pedicels

prescent.

ascending to spreading, 1.5–2 mm.

Flowers

bisexual and unisexual, with staminate, pistillate, or sometimes both sexes occurring with bisexual flowers on the same plant, 1–2(–5) per ocreate fascicle, base stipelike;

perianth accrescent, white to greenish white, campanulate, glabrous;

tepals 5, connate proximally, sepaloid, dimorphic, outer slightly larger than inner.

1–2(–5) per ocreate fascicle;

perianth yellowish green or greenish;

tepals connate ca. 1/4 their length, lanceolate-ovate to obovate, 2–4 mm, apex rounded to acute.

Staminate flowers

stamens 8 (9);

filaments distinct, adnate to base of perianth tube, glabrous;

anthers yellow or pink to purple, ovate to elliptic;

pistil rudimentary.

anthers pink to purple, ovate to elliptic.

Pistillate flowers

tube white or reddish purple to black in fruit, becoming fleshy;

stamens rudimentary;

styles 3, spreading, connate proximally;

stigmas fimbriate.

tube white in fruit.

Achenes

completely or partly included in fleshy perianth, black or dark brown, unwinged, 3-gonous to subglobose, glabrous.

exserted or included, black or dark brown, 3-gonous, 3–4 × 2–3 mm, shiny, smooth.

Seeds

embryo straight.

x

= 10.

2n

= 20 (New Zealand).

Muehlenbeckia

Muehlenbeckia complexa

Phenology Flowering Jul–Sep.
Habitat Sunny, disturbed sites, often in urban areas
Elevation 0-500 m (0-1600 ft)
Distribution
from USDA
Central America; South America; Pacific Islands (New Zealand); Australia [Introduced in North America]
[BONAP county map]
from FNA
CA; Pacific Islands (New Zealand) [Introduced in North America]
[WildflowerSearch map]
[BONAP county map]
Discussion

species 23 (2 in the flora)

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Muehlenbeckia complexa is cultivated as an ornamental and escapes rarely in the flora area.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Tubes of pistillate flowers white in fruit; distal branches of stems brownish-puberulent; leaf blades ovate-oblong to suborbiculate or panduriform
M. complexa
1. Tubes of pistillate flowers reddish purple to black in fruit; distal branches of stems glabrous; leaf blades triangular-lanceolate
M. hastulata
Source FNA vol. 5, p. 485. Author: Craig C. Freeman. FNA vol. 5, p. 485.
Parent taxa Polygonaceae > subfam. Polygonoideae Polygonaceae > subfam. Polygonoideae > Muehlenbeckia
Sibling taxa
M. hastulata
Subordinate taxa
M. complexa, M. hastulata
Synonyms Polygonum complexum
Name authority Meisner: Pl. Vasc. Gen. 1: 316 (A. Cunningham) Meisner: Pl. Vasc. Gen. 2: 227. (1841)
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