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horn calcareous moss, lipstick thyme moss

largetooth calcareous moss, Menzies' red-mouth minium, red-mouth leafy moss

Habit Plants 1.5–5(–8) cm. Plants 0.5–1(–2.5) cm.
Stems

red or brown.

red or reddish brown.

Leaves

green, yellowish, or occasionally dark green, slightly contorted, usually coiled and twisted when dry, sometimes to one side of stem, narrowly elliptic, narrowly ovate-elliptic, or ovate-lanceolate, 3–6 mm, proximal stem leaves narrowly triangular;

base short-decurrent;

margins pale brown, 2-stratose, toothed to below mid leaf, sometimes to near base, teeth paired, large, sharp;

apex acute or acuminate, apiculate or cuspidate, cusp sometimes toothed;

costa ending well below apex or occasionally percurrent, distal abaxial surface strongly toothed, adaxial surface occasionally toothed;

medial laminal cells ± isodiametric or short-elongate, (15–)17–26(–30) µm, slightly smaller towards margins, not in diagonal rows, sometimes in longitudinal rows, not or weakly collenchymatous, blue postmortal color absent;

marginal cells linear, in (2–)3–4 rows.

green to dark green, slightly contorted or undulate when dry, obovate or occasionally elliptic, 2.5–5 mm;

base long-decurrent;

margins reddish brown, multistratose with stereid band, toothed distally, teeth paired, long, sharp;

apex acute, obtuse, or rounded-acute, cuspidate, cusp toothed;

costa percurrent or excurrent, distal abaxial surface smooth, sometimes with few small teeth;

medial laminal cells ± isodiametric or short-elongate, 25–40 µm, smaller towards margins, not in rows, not collenchymatous, blue postmortal color absent;

marginal cells linear, in (3–)4–5 rows.

Seta

single.

single or multiple.

Sexual condition

dioicous.

synoicous.

Capsule

pale brown, 3–5 mm;

operculum conic-mammillate, rarely short-rostrate;

exostome yellowish brown.

pale yellow, 1.5–4 mm;

operculum rostrate;

exostome dark reddish brown, often purplish.

Spores

30–35 µm.

16–24 µm.

Mnium hornum

Mnium spinulosum

Phenology Capsules mature summer. Capsules mature late spring.
Habitat Banks along streams, shorelines, cliff faces on moist, sandy soil, humus Forests, on humus, logs, rock, base of trees
Elevation low to moderate elevations low to moderate elevations
Distribution
from FNA
AL; CO; CT; DE; FL; GA; IA; IL; KY; MA; MD; ME; NC; NJ; NY; OH; PA; RI; TN; VA; VT; WV; NB; NL; NS; ON; PE; QC; South America; Asia; Greenland; Europe; Africa
[WildflowerSearch map]
from FNA
AK; AZ; CA; CT; GA; ID; IL; KY; MA; MD; ME; MI; MN; MT; NH; NY; OR; RI; SD; VT; WA; WI; AB; BC; MB; NB; NL; NS; NT; ON; PE; QC; SK; YT; Europe; Asia
[WildflowerSearch map]
Discussion

Mnium hornum is one of the most distinctive members of the genus. The long, narrow leaves and strongly toothed abaxial costae with teeth of 1(–3) cells are diagnostic.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Mnium spinulosum is distinguished by usually obovate leaves with broad, thick, and pigmented margins, and dark reddish brown exostomes. The capsule mouths are dark reddish brown; H. A. Crum and L. E. Anderson (1981) referred to this species as the red-mouthed mnium.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 28, p. 226. FNA vol. 28, p. 228.
Parent taxa Mniaceae > Mnium Mniaceae > Mnium
Sibling taxa
M. arizonicum, M. blyttii, M. lycopodioides, M. marginatum, M. spinosum, M. spinulosum, M. stellare, M. thomsonii
M. arizonicum, M. blyttii, M. hornum, M. lycopodioides, M. marginatum, M. spinosum, M. stellare, M. thomsonii
Name authority Hedwig: Sp. Musc. Frond., 188. (1801) Bruch & Schimper: Bryol. Europ. 4: 206, plate 394. (1846)
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