Mirabilis nyctaginea |
Mirabilis gigantea |
|
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four-o'clock, heart-leaf four-o'clock, heart-leaf umbrella-wort, heartleaf umbrellawort four-o'clock, umbrellawort, wild four-o'clock |
giant four o'clock |
|
Stems | usually erect or ascending, occasionally decumbent, leafy mostly in proximal 2/3 of plant, openly forked distally, 4–15 dm, basally usually glabrous or puberulent in 2 lines, rarely spreading-pubescent; distally stems usually puberulent in 2 lines, occasionally glabrate, rarely spreading glandular-pubescent. |
erect, moderately leafy in proximal 2/3 of plant, few branched distally, 5–20 dm, basally finely pubescent with appressed, curled hairs often in 2 bands or rarely glabrate; distally densely puberulent with appressed, curled hairs. |
Leaves | ascending at 45–80°, abruptly reduced to inflorescence; petiole 0.2–2 cm; blade green, ovate-lanceolate to ovate or triangular, 3–10 × 2–6.5 cm, usually ± thin, base obtuse, round, truncate, or cordate, apex acute to acuminate, rarely rounded, surfaces usually glabrous, sometimes puberulent or sparsely hispidulous. |
ascending at 30–45°, progressively reduced toward infloresence; petiole 0.2–1 cm; blade grayish green, ovate to broadly lanceolate or oblong, 5–10.5 × 1.8–8 cm, coriaceous, base cuneate to round, truncate, or cordate, apex usually acute to obtuse, occasionally round, surfaces usually glabrous, sometimes very sparsely puberulent with curved hairs. |
Inflorescences | terminal and in upper axils, subumbellate clusters at ends of long, forked branches; peduncle 5–20 mm, usually pubescent with ascending, often curved, glandular or eglandular hairs, crosswalls of hairs pale; involucres pale green, often tinged pinkish, widely bell-shaped to almost rotate, 4–6 mm in flower, 8–15 mm in fruit, glabrous or glabrate but with minute curved hairs on margins, or rarely puberulent or pilose throughout, 50–90% connate, lobes ovate to broadly ovate. |
usually terminal, main axis with short side branches, sometimes short branches present in upper axils; peduncle 5–19 mm, usually densely puberulent with curled hairs, sometimes sparsely so, occasionally spreading-villous, crosswalls of hairs pale; involucres pale green, widely bell-shaped, 4–5 mm in flower, 7–9 mm in fruit, sparsely to densely puberulent with curved hairs, 50–70% connate, lobes ovate. |
Flowers | (2–)3(–5) per involucre; perianth usually pink to reddish purple, rarely white, 1 cm. |
3 per involucre; perianth rose-pink to pale pink, 1 cm. |
Fruits | dark grayish brown to reddish brown (ribs and tubercles usually slightly paler), narrowly obovate and tapering at both ends, 3.4–5 mm, shaggy-pubescent with spreading, sometimes tufted, hairs, 0.3–0.4 mm, sometimes also with layer of minute hairs; ribs usually irregularly and deeply notched, especially toward apex, round to bluntly angled, 0.5–0.75 times width of sulci, 0.5–1 times as wide as high; sulci with pale small to tall tubercles that are sometimes horizontally lengthened and shelflike. |
dark grayish brown, narrowly obovoid, 4–5.3 mm, shaggy-pubescent with spreading hairs often tufted on tubercles, hairs 0.3–0.5 mm, also with layer of minute hairs; ribs usually paler, round-angular, 0.5–1 times width of sulci, as wide as high, irregularly notched or tuberculate, often most deeply so near apex; sulci usually with small dark tubercules or raised, horizontal, riblike tubercles. |
2n | = 58. |
|
Mirabilis nyctaginea |
Mirabilis gigantea |
|
Phenology | Flowering late spring–early fall. | Flowering spring–fall. |
Habitat | Weedy areas in dry, often disturbed sites | Sandy, open fields or woods |
Elevation | 100-2200 m (300-7200 ft) | 200-300 m (700-1000 ft) |
Distribution |
AL; AR; AZ; CA; CO; CT; DC; DE; IA; ID; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; MT; NC; ND; NE; NH; NJ; NM; NV; NY; OH; OK; OR; PA; RI; SD; TN; TX; UT; VA; VT; WA; WI; WV; WY; AB; BC; MB; ON; QC; SK; Europe [Introduced in Mexico]
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OK; TX |
Discussion | Mirabilis nyctaginea is considered a noxious weed in some states. The holotype of Mirabilis ×collina Shinners is a hybrid between M. nyctaginea and M. albida. On the Great Plains, M. nyctaginea also appears to intergrade with M. albida. Prominence of the tubercles and redness of the fruits decreases in western populations. Near the Great Lakes, comparatively narrow-leaved plants with sparsely hirsute stems seem to be intergrades between M. nyctaginea and more or less hirsute M. albida. Mirabilis ×serotina Shinners is a hybrid between M. nyctaginea and M. glabra. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
In the northern portion of the range, Mirabilis gigantea appears to intergrade with M. albida. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 4, p. 55. | FNA vol. 4, p. 55. |
Parent taxa | Nyctaginaceae > Mirabilis > sect. Oxybaphus | Nyctaginaceae > Mirabilis > sect. Oxybaphus |
Sibling taxa | ||
Synonyms | Allionia nyctaginea, Oxybaphus nyctagineus | Allionia gigantea, Oxybaphus giganteus |
Name authority | (Michaux) MacMillan: Metasp. Minnesota Valley, 217. (1892) | (Standley) Shinners: Field & Lab. 19: 177. (1951) |
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